PBDB Taxon

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Nyssaceae

Angiospermae - Cornales - Nyssaceae

Taxonomy

Nyssaceae is a family. It is extant.

It was assigned to Myrtiflorae by MacGinitie (1941); to Dicotyledones by Chandler (1961), Chandler (1962) and Chandler (1963); and to Cornales by The Angiosperm Phylogeny Group (2016).

Subtaxa

Camptotheca, Davidoidea, Lucknowia, Nyssa, Nyssites, Nyssoidea, Nyssoxylon, Palaeonyssa, Paleodavidia, Protonyssa, Pseudonyssa, Tsukada

Synonyms

Davidiaceae is a family. It is extant. It was considered monophyletic by The Angiosperm Phylogeny Group (2016).

It was synonymized subjectively with Nyssaceae by The Angiosperm Phylogeny Group (2016).

It was assigned to Cornales by Wolfe and Wehr (1987).

Synonymy list

p. 21

Year	Name and author
1941	Nyssaceae MacGinitie
1961	Nyssaceae Chandler p. 274
1962	Nyssaceae Chandler p. 120
1963	Nyssaceae Chandler p. 123
1987	Davidiaceae Wolfe and Wehr p. 21
2016	Nyssaceae The Angiosperm Phylogeny Group pp. 8, 19

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Rank	Name	Author
kingdom	Plantae
phylum	Spermatophyta
class	Angiospermae
—	Mesangiosperms
—	Eudicots	Doyle and Hotton 1991
—	Coreeudicots	The Angiosperm Phylogeny Group 2009

Rank	Name	Author
—	Superasterids
—	Asterids
order	Cornales	Link 1829
family	Nyssaceae
family	Nyssaceae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Nyssaceae

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G. Camptotheca Decaisne 1873

†Camptotheca kyushuensis Suzuki 1975

G. †Davidoidea Johnson 1937

†Davidoidea hebridica Johnson 1937

G. †Lucknowia Thiergart and Frantz 1963

†Lucknowia sahnii Thiergart and Frantz 1963

G. Nyssa Linnaeus 1753 [tupelo]

†Nyssa acuticostata Perkins 1906

Nyssa aquatica Linnaeus 1753

†Nyssa aquaticaformis Berry 1916

†Nyssa arctica Heer 1869

†Nyssa ascoidea Perkins 1904

Nyssa biflora Walter 1788

†Nyssa buddiana Ward 1885

†Nyssa californica MacGinitie 1941

†Nyssa clarkii Perkins 1904

†Nyssa complanata Lesquereux 1861

†Nyssa cooperi Chandler 1961

†Nyssa crassicostata Perkins 1904

†Nyssa cuneata Newberry 1863

†Nyssa curta Perkins 1904

†Nyssa cylindrica Perkins 1904

†Nyssa denveriana Knowlton 1919

†Nyssa disseminata Kirchheimer 1937

†Nyssa elongata Perkins 1904

†Nyssa eolignitica Berry 1916

†Nyssa equicostata Perkins 1904

†Nyssa europaea Unger 1845

†Nyssa excavata Perkins 1904

†Nyssa gracilis Berry 1909

†Nyssa jonesii Perkins 1904

†Nyssa laevigata Lesquereux 1861

†Nyssa lamellosa Perkins 1904

†Nyssa lanceolata Lesquereux 1872

†Nyssa lescurii Hitchcock 1862

†Nyssa microcarpa Lesquereux 1861

†Nyssa multicostata Perkins 1904

†Nyssa ovalis Perkins 1906

†Nyssa ovata Perkins 1904

†Nyssa racemosa Knowlton 1898

†Nyssa snowiana Lesquereux 1891

†Nyssa solea Perkins 1904

Nyssa sylvatica Marshall 1785

Nyssa uniflora Walter 1788

†Nyssa vetusta Newberry 1868

Invalid names: Magnolia obovata Newberry 1868 [synonym]

†Nyssa wilcoxiana Berry 1916

G. †Nyssites Geyler 1887

†Nyssites obovatus Weber 1851

G. †Nyssoidea Chandler 1962

†Nyssoidea eocenica Chandler 1962

G. †Nyssoxylon Maedel 1959

†Nyssoxylon japonicum Maedel 1959

G. †Palaeonyssa Reid and Chandler 1933

†Palaeonyssa multilocularis Reid and Chandler 1933

G. †Paleodavidia Miki 1956

†Paleodavidia multipterium Miki 1941

G. †Protonyssa Reid and Chandler 1933

†Protonyssa bilocularis Reid and Chandler 1933

G. †Pseudonyssa Kinkelin 1900

†Pseudonyssa palmiformis Kinkelin 1900

Invalid names: Davidiaceae [synonym]

Diagnosis

Reference	Diagnosis
The Angiosperm Phylogeny Group 2016	Nyssaceae, included in Cornaceae in APG III (2009), have been shown by molecular studies (Xiang et al., 2011) to include Camptothecaceae, Davidiaceae and Mastixiaceae, which are sister to a clade comprising Hydrostachyaceae, Loasaceae and Hydrangeaceae. They are therefore widely separated from Cornaceae, and thus Nyssaceae need to be accepted.