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Harpetidae

Artiopoda - Harpetida - Harpetidae

Synonyms
Synonymy list
YearName and author
1847Harpidae Hawle and Corda
1847Harpetidae Hawle and Corda pp. 1-176
1854Arraphidae Angelin
1982Harpidae Romano and Henry p. 625
1998Harpetidae Sepkoski
2002Harpetidae Ebach and McNamara
2009Harpetidae McNamara et al. p. 14
2011Harpetidae Adrain p. 107
2021Harpetidae Beech and Lamsdell p. 2221

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
RankNameAuthor
phylumArthropodaGravenhorst 1843
classArtiopodaHou and Bergstrom
classTrilobitaWalch 1771
orderHarpetida(Whittington 1959)
familyHarpetidaeHawle and Corda 1847
familyHarpetidaeHawle and Corda 1847

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Harpetidae Hawle and Corda 1847
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G. †Bohemoharpes Vanek 1963
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Subg. †Bohemoharpes (Bohemoharpes) Vanek 1963
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†Bohemoharpes (Bohemoharpes) hypsipyle Pribyl and Vanek 1986
Subg. †Bohemoharpes (Unguloharpes) Pribyl and Vanek 1981
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†Bohemoharpes bubovicensis Pribyl and Vanek 1986
†Bohemoharpes hypsipyle Pribyl and Vanek 1986
†Bohemoharpes inflexa Doubrava 1991
†Bohemoharpes janae Doubrava 1991
†Bohemoharpes naumanni Barrande 1852
†Bohemoharpes ovatus Boucek 1935
†Bohemoharpes ungula Sternberg 1833
†Bohemoharpes vittatus Barrande 1852
†Bohemoharpes wilkensii Münster 1840
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Invalid names: Harpes crassifrons Barrande 1846 [synonym]
Invalid names: Declivolithus Pribyl and Vanek 1967 [synonym]
G. †Declivoharpes Pribyl and Vanek 1981
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†Bohemoharpes (Declivoharpes) praecedens Prantl and Pribyl 1954
G. †Dolichoharpes Whittington 1949
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†Dolichoharpes arctica Whittington 1954
†Dolichoharpes doranni Whittington 1950
†Dolichoharpes reticulata Whittington 1949
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Invalid names: Dolichoharpes procliva Esker, III 1964 [synonym]
†Dolichoharpes uniseralis Raymond 1925
†Dolichoharpes uniserialis Raymond 1925
†Dolichoharpes villosus Koroleva 1978
G. †Dubhglasina Lamont 1948
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†Dubhglasina depressus Harrington and Leanza 1957
†Dubhglasina expansus Jell 1985
†Dubhglasina parvula M'Coy 1851
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Invalid names: Dubhglasina aldonsensis Lamont 1948 [synonym]
Invalid names: Australoharpes Harrington and Leanza 1957 [synonym], Sinoharpes Sheng 1974 [synonym]
G. †Eoharpes Raymond 1905
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†Eoharpes antiquatus Billings 1859
†Eoharpes benignensis Barrande 1872
†Eoharpes cristatus Romano 1975
†Eoharpes guichenensis Henry and Phillipot 1968
†Eoharpes macaoensis Romano and Henry 1982
†Eoharpes primus Barrande 1872
†Eoharpes yijianfangensis Zhou et al. 2014
G. †Eotrinucleus Zhou and Zhang 1978
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†Eotrinucleus antiquus Peng 1992
G. †Eskoharpes McNamara et al. 2009
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†Eskoharpes boltoni McNamara et al. 2009
†Eskoharpes friendi McNamara et al. 2009
†Eskoharpes guthae McNamara et al. 2009
†Eskoharpes neogracilis Richter and Richter 1924
†Eskoharpes palanasus McNamara et al. 2009
†Eskoharpes sicarius Johnson 2024
†Eskoharpes socialis Holzapfel 1895
†Eskoharpes teicherti McNamara et al. 2009
†Eskoharpes wandjina McNamara et al. 2009
Invalid names: Globoharpes McNamara et al. 2009 [synonym]
G. †Fissocephalus Lermontova 1951
G. †Fritchaspis Vanek 1963
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†Fritchaspis edgecombei Johnson 2024
†Fritchaspis montagnei Hawle and Corda 1847
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Invalid names: Harpes perneri Prantl and Pribyl 1954 [synonym]
G. †Harpes Goldfuss 1839
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†Harpes acuminatus Lindström 1885
†Harpes boudibensis Johnson 2024
†Harpes dvorcensis Prantl and Pribyl 1954
†Harpes escoti Bergeron 1887
†Harpes fornicatus Novak 1890
†Harpes forojuliensis Gortani 1909
†Harpes gracilis Münster 1840
†Harpes koeneni Wedekind 1914
†Harpes lahceni Johnson 2024
†Harpes latilimbatus Krause 1894
†Harpes lentigo Johnson 2024
†Harpes macrocephallus Goldfuss 1839
†Harpes macrocephalus Goldfuss 1839
†Harpes nymageenensis Fletcher 1975
†Harpes nymageensis Fletcher 1975
†Harpes ormistoni Pribyl and Vanek 1986
†Harpes oudris Johnson 2024
†Harpes parvulus M'Coy 1851
†Harpes prescheri Johnson 2024
†Harpes pyrenaicus Barrios 1886
†Harpes radians Richter 1963
†Harpes reticulatus Vanek 1963
†Harpes sphenocephalus Johnson 2024
†Harpes supercilium Johnson 2024
†Harpes whidbournei Whittington 1950
Invalid names: Reticuloharpes Vanek 1963 [synonym]
G. †Helioharpes Pribyl and Vanek 1981
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†Helioharpes aougili Johnson 2024
†Helioharpes perradiatus Richter and Richter 1943
†Helioharpes transiens Barrande 1872
G. †Hibbertia Jones and Woodward 1899
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†Hibbertia amibouei Lamont 1948
†Hibbertia anticostiensis Twenhofel 1928
†Hibbertia aodiensis Lee 2013
†Hibbertia balclatchiensis Whittington 1950
†Hibbertia conistonensis Lesperance and Weissenberger 1998
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Invalid names: Paraharpes whittingtoni McNamara 1979 [synonym]
†Hibbertia costatus Angelin 1854
†Hibbertia dalecarlicus Thorslund 1930
†Hibbertia hornei Reed 1914
†Hibbertia inghami Owen 1981
†Hibbertia orbicularis Jones and Woodward 1899
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Invalid names: Harpes flanaganni Portlock 1843 [synonym]
†Hibbertia ottawaensis Billings 1865
†Hibbertia perceensis Kindle 1945
†Hibbertia similis Nikolaisen 1965
†Hibbertia trippi Whittington 1950
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Invalid names: Eoharpes ottawaensis Raymond 1910 [synonym], Harpina antiquatus Billings 1859 [synonym]
†Hibbertia wegelini Angelin 1854
†Hibbertia whittingtoni Tripp 1965
Invalid names: Harpesoides Koroleva 1978 [synonym], Metaharpes Lamont 1948 [synonym], Paraharpes Whittington 1950 [synonym], Platyharpes Whittington 1950 [objective synonym], Thorslundops Pribyl and Vanek 1981 [synonym], Wegelinia Pribyl and Vanek 1981 [synonym]
G. †Kathrynia Westrop 1986
G. †Kielania Vanek 1963
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Subg. †Kielania (Kielania) Vanek 1963
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†Kielania (Kielania) obuti Pribyl and Vanek 1986
†Kielania tumula Johnson 2024
†Kielania waageni Prantl and Pribyl 1954
Invalid names: Lowtheria Prantl and Pribyl 1981 [synonym]
G. †Lioharpes Whittington 1950
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Subg. †Lioharpes (Lioharpes) Pribyl and Vanek 1986
†Lioharpes altaicus Weber 1932
†Lioharpes ammari Johnson 2024
†Lioharpes bischofi Roemer 1852
†Lioharpes crassimargo Novak 1890
†Lioharpes galea Johnson 2024
†Lioharpes hastatus Lütke 1965
†Lioharpes morocconensis Johnson 2024
†Lioharpes polaris Maksimova 1977
†Lioharpes saredra Johnson 2024
†Lioharpes scopulum Johnson 2024
†Lioharpes sculptus Hawle and Corda 1847
†Lioharpes vektori Doubrava 1991
†Lioharpes venetus Gortani 1915
†Lioharpes venulosus Whittington 1950
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Invalid names: Harpes ruderalis Hawle and Corda 1847 [synonym], Lioharpes (Lioharpes) venulosus Pribyl and Vanek 1986 [synonym], Lioharpes klukovicensis Vanek et al. 1992 [synonym]
G. †Maghroharpes Johnson 2024
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†Maghroharpes azmamarensis Johnson 2024
†Maghroharpes forteyi Johnson 2024
†Maghroharpes hammii Johnson 2024
†Maghroharpes ihmadii Johnson 2024
†Maghroharpes oufatenensis Johnson 2024
†Maghroharpes rouvillei Frech 1887
†Maghroharpes terridus Johnson 2024
†Maghroharpes zguidensis Johnson 2024
G. †Palaeoharpes Lu and Qian 1977
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†Palaeoharpes primigenius Lu and Qian 1974
G. †Pinnuloharpes Johnson 2024
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†Pinnuloharpes apteros Johnson 2024
†Pinnuloharpes chaperon Johnson 2024
†Pinnuloharpes hamarlaghdadensis Cronier et al. 2018
†Pinnuloharpes haustrum Johnson 2024
†Pinnuloharpes igaouii Johnson 2024
†Pinnuloharpes segaouii Johnson 2024
†Pinnuloharpes torquis Johnson 2024
G. †Scotoharpes Lamont 1948
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†Scotoharpes aduncus Fortey 1980
†Scotoharpes cassinensis Whitfield 1897
†Scotoharpes consuetus Billings 1863
†Scotoharpes domina Lamont 1948
†Scotoharpes excavatus Linnarsson 1875
†Scotoharpes filiarum Dean 1979
†Scotoharpes fragilis Raymond 1925
†Scotoharpes granti Billings 1865
†Scotoharpes judex Marr and Nicholson 1888
†Scotoharpes latior Poulsen 1934
†Scotoharpes laurei Jell and Stait 1985
†Scotoharpes lauriei Jell and Stait 1985
†Scotoharpes loma Lane 1972
†Scotoharpes molongloensis Chatterton and Campbell 1980
†Scotoharpes pansa Maksimova 1960
†Scotoharpes raaschi Norford 1973
†Scotoharpes rotundus Bohlin 1955
†Scotoharpes sinensis Grabau 1925
†Scotoharpes singularis Whittington 1965
†Scotoharpes sombrero Owen 1981
†Scotoharpes spasskii Eichwald 1840
†Scotoharpes taimyricus Balaschova 1959
†Scotoharpes tatouyangensis Chan and Fang 1960
†Scotoharpes telleri Weller 1907
†Scotoharpes tobulatus Chugaeva 1975
†Scotoharpes trinucleoides Etheridge and Mitchell 1917
†Scotoharpes vetustus Zhou and Zhang 1978
†Scotoharpes vitilis Whittington 1963
†Scotoharpes volsellatus Howells 1982
†Scotoharpes willsi Whittington 1950
†Scotoharpes youngi Reed 1914
Invalid names: Aristoharpes Whittington 1950 [synonym], Selenoharpes Whittington 1950 [synonym]
G. †Stoloharpes Johnson 2024
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†Stoloharpes calceolus Johnson 2024
†Stoloharpes capricornus Johnson 2024
†Stoloharpes retiarius Johnson 2024
†Stoloharpes rissaniensis Johnson 2024
Invalid names: Arraphidae Angelin 1854 [synonym], Harpidae Hawle and Corda 1847 [synonym]
Diagnosis
ReferenceDiagnosis
J. D. Beech and J. C. Lamsdell 2021Eye tubercles each with two lenses; semicircular alae adjacent to posterior glabellar lobes; bilamellar fringe with opposed pits in outer surfaces, genal rolls steeply sloping, brim gently sloping, with stout girder on lower lamella separating these two parts, flattened prolongations of fringe varying in length; cephalic suture skirts marginal band of fringe. Hypostome pear-shaped in outline, with ovate middle body, large anterior, small posterior, and wings. Thorax with 12–29 segments, pleurae bent down ventrally at tips. Pygidium small, short (sag.), triangular, with few segments. External surface of glabella and genae with raised ridges in reticulate pattern, tuberculate, or smooth; minute tubercles on fringe between pits and marginal band, on internal rim, and elsewhere (modified from Fortey & Owens 1997).