Kham-keut, at Kham-mon (Schwagerina princeps horizon) (Permian to of Laos)

Where: Laos (18.3° N, 105.0° E: paleocoordinates 16.7° S, 101.9° E)

• coordinate based on political unit

• local area-level geographic resolution

When: Asselian to Asselian (298.9 - 290.1 Ma)

• Waterhouse (1973) considered the fusulinid and brachiopod fauna to indicate an Asselian age, but the region is faulted and folded, making age determination difficult given the poor geographic resolution of sample. Archbold (1981) mentioned Kham-keut as "Asselian to early Sakmarian."

• group of beds-level stratigraphic resolution

Environment/lithology: lithified limestone

Size class: macrofossils

Collected by Dussault, Roux, Magnin

Primary reference: H. Mansuy. 1913. Faunes des calcaires à Productus de l'Indochine. Première série. Mémoires du Service Géologiques de l'Indochine 2(4):1-133 [P. Wagner/P. Wagner/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 233571: authorized by Matthew Clapham, entered by Matthew Clapham on 18.02.2024

Creative Commons license: CC BY (attribution)

Taxonomic list

Rhynchonellata
 Spiriferida - Reticulariidae
"Reticularia" contexta n. sp. Mansuy 1913
 Spiriferida - Elythidae
""Reticularia" ? rostrata" = Bajkuria rostrata Kutorga 1842
 Spiriferida - Paeckelmanellidae
""Spirifer" ? dieneri" = Paeckelmanella dieneri
""Spirifer" ? dieneri" = Paeckelmanella dieneri Tschernyschew 1902
 Spiriferida - Strophopleuridae
""Spiriferina" (Spiriferella) artiensis" = Spiriferinaella artiensis, ""Spirifer" ? tastubensis" = Spiriferinaella tastubensis
""Spiriferina" (Spiriferella) artiensis" = Spiriferinaella artiensis Stuckenberg 1898
mut. orientalis
""Spirifer" ? tastubensis" = Spiriferinaella tastubensis Tschernyschew 1902
 Spiriferida - Ambocoeliidae
Ambocoelia cf. urii, "Ambocoelia planoconvexa" = Crurithyris planoconvexa
Ambocoelia cf. urii Fleming 1823
"Ambocoelia planoconvexa" = Crurithyris planoconvexa Shumard 1855
 Spiriferida - Choristitidae
""Spirifer" ? trigonalis" = Brachythyrina rectangula, ""Spirifer" ? fritschi" = Choristites fritschi
""Spirifer" ? trigonalis" = Brachythyrina rectangula Kutorga 1844
""Spirifer" ? fritschi" = Choristites fritschi Schellwien 1892
 Spiriferida - Spiriferidae
"Spirifer" ? uralicus, ""Spirifer" ? lyra" = Eliva lyra, ""Spirifer" ? panduriformis" = Eliva panduriformis, ""Spirifer" ? nikitini" = Purdonella nikitini
"Spirifer" ? uralicus Tschernyschew 1902
""Spirifer" ? lyra" = Eliva lyra Kutorga 1844
""Spirifer" ? panduriformis" = Eliva panduriformis Kutorga 1844
""Spirifer" ? nikitini" = Purdonella nikitini Tschernyschew 1902
 Spiriferida - Trigonotretidae
""Spirifer" cf. marcoui" = Betaneospirifer marcoui, ""Spirifer" ? musakheylensis" = Betaneospirifer moosakhailensis, ""Spirifer" ? cameratus" = Neospirifer cameratus
""Spirifer" cf. marcoui" = Betaneospirifer marcoui Waagen 1883
""Spirifer" ? musakheylensis" = Betaneospirifer moosakhailensis Davidson 1862
""Spirifer" ? cameratus" = Neospirifer cameratus Morton 1836
 Spiriferida - Spiriferellidae
""Spirifer" ? tibetanus" = Elivina tibetana, ""Spirifer" ? interplicatus" = Spiriferella interplicata
""Spirifer" ? tibetanus" = Elivina tibetana Diener 1897
""Spirifer" ? interplicatus" = Spiriferella interplicata Rothpletz 1892
 Spiriferida - Brachythyrididae
""Spirifer" ? quadriradiatus" = Brachythyris quadriradiatus
""Spirifer" ? quadriradiatus" = Brachythyris quadriradiatus de Verneuil 1845
 Spiriferida - Martiniidae
""Martiniopsis" ? baschkirica" = Elivella baschkirica, ""Spirifer" ? sokolovi" = Jilinmartinia sokolovi, "Martinia" ? corculum, "Martinia" ? triquetra
""Martiniopsis" ? baschkirica" = Elivella baschkirica Tschernyschew 1902
includes mut. indosinensis
""Spirifer" ? sokolovi" = Jilinmartinia sokolovi Tschernyschew 1902
"Martinia" ? corculum Kutorga 1844
"Martinia" ? triquetra Gemmellaro 1899
 Spiriferida - Ingelarellidae
"Martiniopsis" ? uralica Tschernyschew 1902
"Martiniopsis" ? orientalis Tschernyschew 1902
 Terebratulida - Notothyrididae
""Notothyris" (Rostranteris) ? mediterranea" = Rostranteris mediterraneum
""Notothyris" (Rostranteris) ? mediterranea" = Rostranteris mediterraneum Gemmellaro 1899
mut. laosensis
 Terebratulida - Gillediidae
""Dielasma" ? juresanense" = Pyandzhelasma juresanense
""Dielasma" ? juresanense" = Pyandzhelasma juresanense Tschernyschew 1902
 Terebratulida - Heterelasminidae
""Hemiptychina" ? dieneri" = Heterelasmina dieneri
""Hemiptychina" ? dieneri" = Heterelasmina dieneri Gemmellaro 1899
 Terebratulida - Dielasmatidae
"Dielasma" ? vesiculare de Koninck 1851
"Dielasma" ? moelleri Tschernyschew 1902
 Athyridida - Athyrididae
""Athyris" ? gerardi" = Himathyris gerardi Diener 1899
"Athyris" cf. lamellosa Léveillé 1835
 Spiriferinida - Paraspiriferinidae
""Spiriferina" ? ornata" = Callispirina ornata, "Spiriferina multipunctata n. sp." = Callispirina multipunctata
""Spiriferina" ? ornata" = Callispirina ornata Waagen 1883
"Spiriferina multipunctata n. sp." = Callispirina multipunctata Mansuy 1913
 Spiriferinida - Spiriferellinidae
"Spiriferina rouxi n. sp." = Spiriferellina rouxi
"Spiriferina rouxi n. sp." = Spiriferellina rouxi Mansuy 1913
 Spiriferinida - Spiriferinidae
"Spiriferina" triplicata n. sp. Mansuy 1913
 Spiriferinida - Licharewiidae
""Spirifer" cf. curvirostris" = Licharewia curvirostris
""Spirifer" cf. curvirostris" = Licharewia curvirostris de Verneuil 1845
 Orthida - Rhipidomellidae
""Orthis" (Schizophoria) ? michelini" = Rhipidomella michelini
""Orthis" (Schizophoria) ? michelini" = Rhipidomella michelini L'Eveille 1835
mut. orientalis
 Orthida - Schizophoriidae
""Orthis" (Schizophoria) ? supracarbonica" = Schizophoria supracarbonica
""Orthis" (Schizophoria) ? supracarbonica" = Schizophoria supracarbonica Tschernyschew 1902
 Orthida - Enteletidae
Enteletes khammonensis n. sp., "Enteletes mirabilis n. sp." = Enteletina mirabilis
Enteletes khammonensis n. sp. Mansuy 1913
"Enteletes mirabilis n. sp." = Enteletina mirabilis Mansuy 1913
 Rhynchonellida -
Laosia dussaulti n. gen. n. sp.
Laosia dussaulti n. gen. n. sp. Mansuy 1913
 Rhynchonellida - Pugnacidae
"Pugnax" ? osagensis Swallow 1908
"Pugnax" sublaevis n. sp. Mansuy 1913
 Rhynchonellida - Stenoscismatidae
""Camarophoria" ? mutabilis" = Stenoscisma mutabilis, "Camarophoria" ? crumena
""Camarophoria" ? mutabilis" = Stenoscisma mutabilis Tschernyschew 1902
 Rhynchonellida - Wellerellidae
"Uncinulus timorensis" = Uncinunellina timorensis
"Uncinulus timorensis" = Uncinunellina timorensis Beyrich 1865
mut. laosensis
Strophomenata
 Strophomenida - Rugosochonetidae
""Chonetes" ? variolata" = Neochonetes variolata
""Chonetes" ? variolata" = Neochonetes variolata d'Orbigny 1842
 Orthotetida - Meekellidae
Meekella eximia Eichwald 1845
 Productida - Aulostegidae
Urushtenia sp.1 Licharew 1935
 Productida - Echinoconchidae
Karavankina sp.2, ""Productus" ? pustulosus" = Pustula pustulosa, ""Productus" scalaris n. sp." = Productus (Juresania) scalaris
Karavankina sp.2 Ramovs 1966
""Productus" ? pustulosus" = Pustula pustulosa Phillips 1836
""Productus" scalaris n. sp." = Productus (Juresania) scalaris Mansuy 1913
 Productida - Linoproductidae
Linoproductus cambodgensis3, ""Productus" ? tenuistriatus" = Linoproductus tenuistriatus, ""Productus" ? konincki" = Cancrinella koninckiana, ""Productus" ? cancriniformis" = Cancrinella cancriniformis
Linoproductus cambodgensis3 Termier and Termier 1970
""Productus" ? tenuistriatus" = Linoproductus tenuistriatus de Verneuil 1845
""Productus" ? konincki" = Cancrinella koninckiana Keyserling 1846
In text "probably higher than S. princeps" but S. princeps in table
""Productus" ? cancriniformis" = Cancrinella cancriniformis Tschernyschew 1889
 Productida - Monticuliferidae
"Proboscidella" avellana n. sp. Mansuy 1913
 Productida - Paucispiniferidae
"Productus margaritatus n. sp." = Transennatia margaritatus
"Productus margaritatus n. sp." = Transennatia margaritatus Mansuy 1913
 Productida -
? Marginifera depressa n. sp. Mansuy 1913
Marginifera involuta Tschernyschew 1902
Marginifera gibbosa n. sp. Mansuy 1913
 Productida - Productellidae
""Productus" ? pustulatus" = Krotovia pustulata, ""Productus" ? iakovlevi" = Krotovia yakovlevi, "Productus dussaulti n. sp." = Stictozoster dussaulti, "Productus propinquus n. sp." = Stictozoster propinquus, ""Proboscidella" ? kutorgae" = Rugivestis kutorgae, "Productus" ? rectangulus, "Productus" radula n. sp., "Productus" acutauritus n. sp., "Productus" cf. plicatilis, "Productus" concinnus n. sp., ""Productus" ? boliviensis" = Alpavlia boliviensis, ""Productus" ? gruenewaldti" = Chaoiella gruenewaldti
""Productus" ? pustulatus" = Krotovia pustulata Keyserling 1853
""Productus" ? iakovlevi" = Krotovia yakovlevi Tschernyschew 1902
var. laosensis
"Productus dussaulti n. sp." = Stictozoster dussaulti Mansuy 1913
"Productus propinquus n. sp." = Stictozoster propinquus Mansuy 1913
""Proboscidella" ? kutorgae" = Rugivestis kutorgae Tschernyschew 1902
"Productus" ? rectangulus
mut. triplicatus
"Productus" radula n. sp. Mansuy 1913
"Productus" acutauritus n. sp. Mansuy 1913
"Productus" concinnus n. sp. Mansuy 1913
""Productus" ? boliviensis" = Alpavlia boliviensis d'Orbigny 1842
""Productus" ? gruenewaldti" = Chaoiella gruenewaldti Krotov 1888
 Productida - Productidae
""Productus" ? porrectus" = Kochiproductus porrectus, ""Productus" ? timanicus" = Sowerbina timanica, ""Productus" ? transversalis" = Reticulatia transversalis
""Productus" ? porrectus" = Kochiproductus porrectus Kutorga 1844
""Productus" ? timanicus" = Sowerbina timanica Stuckenberg 1875
""Productus" ? transversalis" = Reticulatia transversalis Tschernyschew 1902
Stenolaemata
 Cryptostomata - Acanthocladiidae
Acanthocladia cf. anceps, "? Synocladia cf. virgulacea" = Synocladia dubia, Polypora sp.
Acanthocladia cf. anceps Schlotheim 1820
"? Synocladia cf. virgulacea" = Synocladia dubia Schlotheim 1820
 Cryptostomata - Fenestellidae
Fenestella laosensis n. sp. Mansuy 1913
Gastropoda
 Ptychomphaloidea - Ptychomphalidae
Ptychomphalus venustus n. sp. Mansuy 1913 snail
 Murchisoniina - Eotomariidae
Euconospira striatula Mansuy 1913 snail
 Cycloneritida - Trachyspiridae
Trachydomia dussaulti n. sp. Mansuy 1913 snail
 Cyrtoneritida - Orthonychiidae
"Platyceras gracile n. sp." = Orthonychia gracilis
"Platyceras gracile n. sp." = Orthonychia gracilis Mansuy 1913 snail
 Bellerophontida - Bellerophontidae
"Bellerophon cf. jonesianus" = Waagenella jonesianus
"Bellerophon cf. jonesianus" = Waagenella jonesianus de Koninck 1863 snail
 Euomphalina - Platyceratidae
Platyceras khammonense n. sp. Mansuy 1913 snail
Bivalvia
 Pectinida - Streblochondriidae
Streblopteria magnini Mansuy 1913 scallop
 Pectinida - Aviculopectinidae
Aviculopecten laosensis n. sp. Mansuy 1913 scallop
Trilobita
 Proetida - Phillipsiidae
"Phillipsia" proetoides n. sp. Mansuy 1913 trilobite