Shanwang, Linqu County, collection 79/80 (Beijing Museum): Burdigalian, China

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Diptera - Tipulidae
Tipula insolita n. sp. Zhang 1989
Zhang 1989 1 specimen
79027 (2 measurements)
Insecta - Diptera - Bibionidae
Plecia ludongensis n. sp. Hong and Wang 1985
Hong and Wang 1985 1 specimen
80170 (1 measurement)
Plecia solaris n. sp. Zhang 1989
Zhang 1989 1 specimen
79125/6 (Bibio shanwangensis in Hong, 1985 - fig 26) (2 measurements)
Clothonopsis miocenica n. gen., n. sp. Hong and Wang 1987
Hong and Wang 1987 1 specimen
nomen dubium belonging to Plecia
BNHM 80195 (2 measurements)
Bibio cratopodus Zhang 1989
Zhang 1989 1 specimen
79092
Bibio miocenica n. sp. Hong and Wang 1985
Hong and Wang 1985 1 specimen
corrected as Bibio miocenicus
80117 (2 measurements)
Fungivora bullata n. sp. Hong 1985
Zhang 1989 1 specimen
recombined as Bibio bullatus
79003 (2 measurements)
Bibio ventricosus n. sp. Zhang 1989
Zhang 1989 1 specimen
79033 (Bibio shanwangensis in Hong, 1985 - pl 7, figs 1, 2, 3) (2 measurements)
Bibio brunneus n. sp. Zhang 1989
Zhang 1989 1 specimen
79120 (Bibio sp. in Hong, 1985) (2 measurements)
Bibio ruptineurus n. sp. Zhang 1989
Zhang 1989 1 specimen
79123 (Bibio sp. in Hong, 1985) (2 measurements)
Bibio expansus Zhang 1989
Zhang 1989 1 specimen
79053 (Bibio shanwangensis in Hong, 1985 - pl 7, fig 4)
Bibio deformatus n. sp. Zhang 1989
Zhang 1989 1 specimen
79057 (Bibio shanwangensis in Hong, 1985 - pl 4, fig 1) (2 measurements)
Bibio shanwangensis Hong 1984
Hong and Wang 1985 1 specimen
80123
Insecta - Diptera - Sciaridae
Sciara brevipetiola n. sp. Zhang 1989
Zhang 1989 1 specimen
79004 (2 measurements)
Insecta - Hemiptera - Cicadidae
Oncotympana lapidescens n. sp. Zhang 1989
Zhang 1989 1 specimen
recombined as Tanyocicada lapidescens
79023 (1 measurement)
Insecta - Hemiptera - Aphididae
Cinara reconditivenia n. sp. Zhang 1989
Zhang 1989 1 specimen
recombined as Cinara (Cinara) reconditivenia
79056 (2 measurements)
Insecta - Hemiptera
Heteroptera indet. Latreille 1810
Zhang 1989 1 specimen
79086
Insecta - Hemiptera - Urostylididae
Urochela melaina n. sp. Zhang 1989
Zhang 1989 1 specimen
79024 (2 measurements)
Coreiopsis punctatus n. sp. Hong 1983
Zhang 1989 1 specimen
replaced by Urochela maculosa
79084 (2 measurements)
Insecta - Hemiptera - Coreidae
Shanwangicoris longa n. gen., n. sp. Hong and Wang 1987
Hong and Wang 1987 1 specimen
80129 (2 measurements)
Gonocerus shandongensis n. sp. Zhang 1989
Zhang 1989 1 specimen
79025 (3 measurements)
Cloresmus shanwangi n. sp. Zhang 1989
Zhang 1989 1 specimen
79005 (2 measurements)
Coriopsis parva n. sp. (Hong and Wang 1987)
Hong and Wang 1987 1 specimen
misspelling Coreiopsis parva
80115 (2 measurements)
Coriopsis miocenica n. sp. (Hong and Wang 1987)
Hong and Wang 1987 1 specimen
misspelling Coreiopsis miocenica
80104 (1 measurement)
Coreiopsis punctata n. gen., n. sp. Hong 1984
1 specimen
79084
Insecta - Hemiptera - Notonectidae
Tarsonecta mecopoda n. gen., n. sp. Zhang 1989
Zhang 1989 1 specimen
79013 (1 measurement)
Insecta - Hemiptera - Belostomatidae
Manocerus stagnans n. gen., n. sp. Zhang 1989
Zhang 1989 1 specimen
79016 (1 measurement)
Insecta - Hemiptera - Nepidae
Coroides xiejiaheense n. gen., n. sp. Hong 1984
1 specimen
nomen dubium belonging to Nepariae
79113
Insecta - Hemiptera - Fulgoridae
Aphaena lithoecia n. sp. Zhang 1989
Zhang 1989 1 specimen
79022 (2 measurements)
Fulgoropsis fusca n. gen., n. sp. Hong 1984
1 specimen
synonym of Limois shanwangensis
79080
Oxycephala xiejiaheensis Hong 1984
Zhang 1989 3 specimens
synonym of Limois shanwangensis
79026, 79079, 79080
Insecta - Hymenoptera - Xyelidae
Megaxyela xiejiaheensis n. sp. Hong 1984
1 specimen
recombined as Xyelecia xiejiaheensis
79048 (3 measurements)
Insecta - Hymenoptera - Cimbicidae
Procimbex shanwangensis n. sp. Hong 1984
1 specimen
recombined as Abia shanwangensis
79038 (3 measurements)
Abia paurocephala n. sp. Zhang 1989
Zhang 1989 1 specimen
79085 (2 measurements)
Procimbex shandongense n. sp. Hong and Wang 1985
Hong and Wang 1985 1 specimen
recombined as Abia shandongensis
80193 (2 measurements)
Insecta - Hymenoptera - Ichneumonidae
Luia lacaris n. gen., n. sp. Hong 1985
Zhang 1989 1 specimen
79037 (3 measurements)
Insecta - Hymenoptera - Thynnidae
Elis pyrula n. sp. Zhang 1989
Zhang 1989 1 specimen
79089 (2 measurements)
Insecta - Hymenoptera - Pemphredonidae
Sinostigma spinalata n. sp. Hong 1984
1 specimen
recombined as Pemphredon spinalatum
79091 (2 measurements)
Insecta - Hymenoptera - Apidae
Apis miocenica n. sp. Hong 1984
3 specimens
79040, 79050; 80133 in Hong & Wang 1985 (2 measurements)
Insecta - Hymenoptera - Formicidae
Paraphaenogaster shanwangensis n. sp. Hong 1984
1 specimen
79047 (3 measurements)
Heteromyrmex atopogaster n. gen., n. sp. Zhang 1989
Zhang 1989 1 specimen
recombined as Zhangidris atopogaster
79054 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 3) (2 measurements)
Rabidia xiejiaheensis n. sp. Hong 1984
1 specimen
recombined as Camponotites xiejiaheensis
79069 (2 measurements)
Camponotus longus n. sp. Zhang 1989
Zhang 1989 1 specimen
recombined as Camponotites longus
79052 (2 measurements)
Camponotus curviansatus n. sp. Zhang 1989
Zhang 1989 1 specimen
recombined as Camponotites curviansatus
79042 (Camponotus shanwangensis in Hong, 1985 - pl 27, fig 1) (2 measurements)
Camponotus shanwangense Hong 1984
33 specimens
recombined as Camponotites shanwangensis
79031; 79043, 79076 (Liometopum jiaoyanshanense in Hong, 1985); ten specimens including 80179 in Hong & Wang 1985
Camponotus microthoracus n. sp. Zhang 1989
Zhang 1989 3 specimens
recombined as Camponotites microthoracus
79046, 79127, 79128 (2 measurements)
Lasius validus n. sp. Zhang 1989
Zhang 1989 1 specimen
79121 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 4) (2 measurements)
? Lasius truncatus n. sp. Zhang 1989
Zhang 1989 1 specimen
79055 (1 measurement)
Lasius vetulus Dlussky 1981
Zhang 1989 1 specimen
79030 (2 measurements)
Camponotus mitis Smith 1892
Zhang 1989 1 specimen
79067 (Apis miocenica in Hing, 1985 - pl 22, fig 3)
Formica linquensis n. sp. Zhang 1989
Zhang 1989 1 specimen
79058 (Paraphaenogaster shanwangense in Hong, 1985 - pl 23, fig 4) (2 measurements)
Liometopum jiaoyanshanense n. sp. Hong 1985
Zhang 1989 2 specimens
recombined as Dolichoderus jiaoyanshanensis
79087; 80196 in Hong & Wang 1985 (4 measurements)
Shanwangella palaeoptera n. gen., n. sp. Zhang 1989
Zhang 1989 1 specimen
recombined as Liometopum palaeopterum
79051 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 2) (2 measurements)
Liometopum potamophilum n. sp. Zhang 1989
Zhang 1989 1 specimen
79036 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 6) (2 measurements)
? Tapinoma baculum n. sp. Zhang 1989
Zhang 1989 1 specimen
79051 (2 measurements)
Insecta - Hymenoptera - Tiphiidae
Brachyopteris shanwangensis n. sp. Hong 1984
1 specimen
recombined as Tiphia shanwangensis
79041 (2 measurements)
Insecta - Hymenoptera - Vespidae
Pseudoperla shanwangensis n. sp. Hong 1984
1 specimen
recombined as Eumenes shanwangensis
79044 (2 measurements)
Insecta - Coleoptera - Carabidae
Shanwangicarabus furvus n. sp. Hong 1984
Hong and Wang 1986 4 specimens
recombined as Chlaenius furvus
80125, 80106, 80126, 480183 (2 measurements)
Shanwangicarabus paucinus n. sp. (Hong and Wang 1985)
Hong and Wang 1986 1 specimen
synonym of Chlaenius furvus
80131 (2 measurements)
? Carabus ovalus n. sp. Hong and Wang 1986
Hong and Wang 1986 1 specimen
80192 (2 measurements)
Calosoma cf. maderae (Fabricius 1775)
Zhang 1989 3 specimens
original and current combination Carabus maderae
79001, 79002, 79108
Scarites mancus n. sp. Zhang 1989
Zhang 1989 1 specimen
79106 (Aliscarabaeus granulatus in Hong, 1985) (1 measurement)
Insecta - Coleoptera - Cupedidae
Cupes longus n. sp. Hong and Wang 1987
Hong and Wang 1987 1 specimen
BNHM 80109 (2 measurements)
Insecta - Coleoptera
Magniscarabaeus furvus Hong 1984
1 specimen
79063 (1 measurement)
    = Polyphaga indet. Emery 1886
Zhang 1989
Insecta - Coleoptera - Endomychidae
Engonius alviolatus (Hong 1984)
Zhang 1989 1 specimen
recombined as Gerstaeckerus alviolatus
79041
Insecta - Coleoptera - Cerambycidae
Cerambycidae informal indet. 1 (Latreille 1802)
Zhang 1989 1 specimen
Cerambicini
79020
Alifasciola daedalea n. sp. Zhang 1989
Zhang 1989 1 specimen
79074 (1 measurement)
Mesosa soteria n. sp. Zhang 1989
Zhang 1989 1 specimen
79008 (1 measurement)
Sinocalosoma longicornuta n. sp. Hong and Wang 1986
Hong and Wang 1986 1 specimen
80175 (1 measurement)
Mesocacia pulla n. sp. Zhang 1989
Zhang 1989 1 specimen
79019 (1 measurement)
Insecta - Coleoptera - Brentidae
Longifistulia parva n. gen., n. sp. Hong and Wang 1987
Hong and Wang 1987 3 specimens
80116 (2 measurements)
Insecta - Coleoptera - Curculionidae
Sinocurculia shanwangensis n. sp. Hong 1985
Hong and Wang 1986 6 specimens
recombined as Atactogaster shanwangensis
80128, 78001, 80107, 80134, 80173, 80174
Sinocurculia granulata n. sp. Hong 1984
1 specimen
recombined as Atactogaster granulata
79109 (1 measurement)
Hylobius aff. niitakensis Kono 1933
Zhang 1989 3 specimens
79106, 79116 (ssp. fukienensis) (1 measurement)
Hylobius medianus n. sp. Zhang 1989
Zhang 1989 1 specimen
79117 (1 measurement)
Hylobius plenus Zhang 1989
Zhang 1989 3 specimens
79010, 79012, 79073
Insecta - Coleoptera - Tenebrionidae
Sinocistela siphla n. gen., n. sp. Zhang 1989
Zhang 1989 1 specimen
79028 (2 measurements)
Insecta - Coleoptera - Meloidae
? Epicauta rufithoraca n. sp. Zhang 1989
Zhang 1989 1 specimen
79072 (1 measurement)
Insecta - Coleoptera
Scarabaeoidea informal indet. 1 Latreille 1802
Zhang 1989 1 specimen
79045 (Magniscarabaeus furvus in Hong & Wang - pl 17, fig 2) (1 measurement)
Insecta - Coleoptera - Geotrupidae
Geotrupidae indet. (Latreille 1802)
Zhang 1989 1 specimen
Geotrupini
79107 (Magniscarabaeus brunneus in Hong, 1985 - pl 20, fig 3) (1 measurement)
Obitiscarabaeus jiaoyanshanense n. gen., n. sp. Hong 1984
1 specimen
recombined as Geotrupes jiaoyanshanense
79114/5 (1 measurement)
Insecta - Coleoptera
Aliscarabaeus granulatus n. sp. Hong 1984
1 specimen
79111
Insecta - Coleoptera - Scarabaeidae
Anomala sp. Samouelle 1819
Zhang 1989 1 specimen
79096
Magniscarabaeus furvus Hong 1984
Hong and Wang 1986 2 specimens
recombined as Anomala furva
80116, 80143
Anomala synemosyna n. sp. Zhang 1989
Zhang 1989 2 specimens
79006, 79007 (1 measurement)
Anomala lochmocola n. sp. Zhang 1989
Zhang 1989 1 specimen
79112 (1 measurement)
Magniscarabaeus brunneus n. gen., n. sp. Hong 1985
Zhang 1989 8 specimens
replaced by Anomala palaeobrunnea
79105; seven including 80101, 80120, 80172, 80188, 80184, 80194, mentioned in Hong & Wang 1987 (2 measurements)
Holotrichia spatha n. sp. Zhang 1989
Zhang 1989 1 specimen
79102 (1 measurement)
Insecta - Coleoptera - Staphylinidae
Lathrobium homorum n. sp. Zhang 1989
Zhang 1989 1 specimen
79021 (1 measurement)
Insecta - Coleoptera - Buprestidae
Procarabus punctatus n. gen., n. sp. Hong 1984
1 specimen
nomen dubium belonging to Eolampra
79118/9 (1 measurement)
Agrilus corrugatus n. sp. Zhang 1989
Zhang 1989 1 specimen
79062 (2 measurements)
Insecta - Coleoptera - Dascillidae
Dascillus shandongianus Zhang 1989
Zhang 1989 4 specimens
79009, 79017, 79077, 79088
Dascillus elongatus Zhang 1989
Zhang 1989 1 specimen
79081
Insecta - Coleoptera
Procarabus longipes n. sp. Hong 1985
Zhang 1989 1 specimen
recombined as Cistelites longipes
79110 (1 measurement)
Insecta - Dermaptera - Forficulidae
Apanechura asecta n. sp. Zhang 1989
Zhang 1989 1 specimen
79029 (1 measurement)
Insecta - Ephemeroptera - Leptophlebiidae
Paraleptophlebia sp. (Lestage 1917)
Zhang 1989 1 specimen
original and current combination Leptophlebia (Paraleptophlebia)
79129 (Baetis sp. in Hong, 1985)
Insecta - Ephemeroptera - Heptageniidae
? Heptagenia sp. Walsh 1863
Zhang 1989 1 specimen
79095
Insecta - Odonata - Gomphaeschnidae
Jagoria sinica n. sp. Zhang 1989
Zhang 1989 1 specimen
79064 (Nasiaeschna orientalis in Hong, 1985) (1 measurement)
Arachnida - Araneae - Sparassidae
Retina robusta Hong 1985
Zhang et al. 1994 1 specimen
recombined as Heteropoda robusta
79104
Arachnida - Araneae - Tetragnathidae
Eocryphoeca parva Hong 1985
Zhang et al. 1994 1 specimen
recombined as Tetragnatha parva
79101
Arachnida - Araneae - Araneidae
Melanites orbiculatus Hong 1985
Zhang et al. 1994 1 specimen
recombined as Tethneus orbiculatus
79100
Araneus ruidipedalis n. sp. Zhang et al. 1994
Zhang et al. 1994 1 specimen
79098
Araneus piceus Lin et al. 1989
Zhang et al. 1994 1 specimen
79099 (part of M. orbiculatus in Hong, 1985)
see common names

Geography
Country:China State/province:Shandong
Coordinates: 36.6° North, 118.7° East (view map)
Paleocoordinates:36.5° North, 116.3° East
Basis of coordinate:based on nearby landmark
Geographic resolution:local area
Time
Period:Neogene Epoch:Miocene
Stage:Burdigalian 10 m.y. bin:Cenozoic 5
Key time interval:Burdigalian
Age range of interval:20.44000 - 15.98000 m.y. ago
Stratigraphy
Formation:Shanwang
Stratigraphic resolution:group of beds
Stratigraphy comments: Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.
Lithology and environment
Primary lithology:diatomaceous,tuffaceous lithified mudstone
Environment:lacustrine - large Tectonic setting:rift
Geology comments: Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.
Taphonomy
Modes of preservation:adpression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Collection method comments: Repository: Beijing Museum of Natural History. This collection contains specimens labeled with five-digit numbers starting with 79... or 80...
Metadata
Database number:128550
Authorizer:M. Clapham, D. Smith Enterer:M. Clapham, E. Leckey
Modifier:M. Clapham Research group:paleoentomology
Created:2012-06-11 02:17:52 Last modified:2017-08-30 23:22:15
Access level:the public Released:2012-06-11 02:17:52
Creative Commons license:CC BY
Reference information

Primary reference:
15208. Y. C. Hong. 1984. Fossil insects in the diatoms of Shanwang. Bulletin of the Tianjin Institute of Geology and Mineral Resources 8:1-15 [D. Smith/E. Leckey/M. Clapham]

Secondary references:

15210 Y. Hong and W. Wang. 1985. Miocene Diptera and Hymenoptera (Insecta) of Shanwang, Shandong Province, China. Memoirs of the Beijing Natural History Museum 31:1-22 [D. Smith/E. Leckey/R. Day]
51641 Y. C. Hong and W. L. Wang. 1986. Miocene Coleoptera (Insecta) of Shanwang, Shandong Province, China. Memoirs of Beijing Natural History Museum 38:1-19 [M. Clapham/M. Clapham]
56339 Y. C. Hong and W. L. Wang. 1987. Miocene Emboptera and Coleoptera (Insecta) of Shanwang, Shandong Province, China. Professional Papers of Stratigraphy and Palaeontology 17:257-262 [M. Clapham/M. Clapham]
51313 Y. C. Hong and W. L. Wang. 1987. Miocene Heteroptera and Coleoptera (Insecta) from Shanwang of Shandong Province, China. Journal of Lanzhou University (Natural Science) 33:116-124 [M. Clapham/M. Clapham]
36942 J. Zhang, B. Sun, and X. Zhang. 1994. Miocene insects and spiders from Shanwang, Shandong 1-298 [M. Clapham/J. Karr]
16579 J. F. Zhang. 1989. Fossil Insects From Shanwang, Shandong, China 1-459 [D. Smith/E. Leckey/M. Clapham]