Khneifiss L2 (Cretaceous to of Syria)

Where: Syria (34.3° N, 37.8° E: paleocoordinates 16.0° N, 32.6° E)

• coordinate estimated from map

When: Late/Upper Campanian to Late/Upper Campanian (83.5 - 66.0 Ma)

Environment/lithology: phosphatic, red, yellow, calcareous phosphorite

• open sea environment
• This phosphatic level is mostly friable, yellow to reddish in colour and very rich in vertebrate remains.

Size class: macrofossils

Primary reference: N. Bardet, H. Cappetta, X. Pereda Suberbiola, M. Mouty, A. K. Al Maleh, A. M. Ahmed, O. Khrata and N. Gannoum. 2000. The marine vertebrate faunas from the Late Cretaceous phosphates of Syria. Geological Magazine 137(3):269-290 [P. Mannion/P. Mannion]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 117852: authorized by Philip Mannion, entered by Philip Mannion on 27.09.2011, edited by Roger Benson

Creative Commons license: CC BY (attribution)

Taxonomic list

• Specimens housed in the Université de Montpellier (UM-Collection Syrie) and Muséum National d’Histoire Naturelle, Paris (MNHN-Collection Syrie)
Chondrichthyes
 Sclerorhynchiformes - Sclerorhynchidae
Ganopristis sp. Arambourg 1935 cartilaginous fish
Ctenopristis sp. Arambourg 1940 cartilaginous fish
 Myliobatiformes - Rhombodontidae
Rhombodus aff. meridionalis Arambourg 1952 stingray
 Rajiformes - Rhinobatidae
Rhinobatos sp. Link 1790 guitarfish
"Rhinobatos sp.1 sp.2"
 Hexanchiformes - Hexanchidae
Hexanchus sp. Rafinesque 1810 sixgill shark
 Lamniformes - Anacoracidae
Squalicorax kaupi Agassiz 1843 crow shark
Squalicorax pristodontus Agassiz 1843 crow shark
 Lamniformes - Odontaspididae
Carcharias sp. Rafinesque 1810 sand shark
 Orectolobiformes - Ginglymostomatidae
Plicatoscyllium sp. Case and Cappetta 1997 nurse shark
 Galeomorphi -
Youssoubatis sp. Cappetta 1992 shark
 Carchariniformes - Scyliorhinidae
Pteroscyllium sp. Cappetta 1980 cat shark
 Galeomorphi -
Hypsobatis sp. Cappetta 1992 shark
 Euselachii - Mitsukurinidae
Anomotodon aff. plicatus Arambourg 1952 goblin shark
Actinopteri
 Teleostei -
Teleostei indet. Müller 1846
 Salmoniformes - Enchodontidae
 Tetraodontiformes - Trigonodontoidae
 Alepisauriformes - Dercetidae
Reptilia
 Testudines -
 Testudines - Cheloniidae
Cheloniidae indet. Bonaparte 1832 sea turtle
 Testudines - Bothremydidae
Taphrosphyini indet. Gaffney et al. 2006 sideneck turtle
 Loricata -
 Squamata - Mosasauridae
Mosasaurinae indet. Gervais 1853 mosasaur
Mosasaurus lemonnieri Dollo 1889 mosasaur
Globidens sp. Gilmore 1912 mosasaur
Prognathodon giganteus Dollo 1889 mosasaur
Plioplatecarpini indet. Russell 1967 mosasaur
"Leiodon anceps" = Liodon anceps Owen 1841 mosasaur
 Sauropterygia - Elasmosauridae
Elasmosauridae indet. Cope 1869 elasmosaur