PBDB Collection

Shanwang, Linqu County, collection 79/80 (Beijing Museum) (Miocene of China)

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone

• Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Beijing Museum of Natural History. This collection contains specimens labeled with five-digit numbers starting with 79... or 80...

Primary reference: Y. C. Hong. 1984. Fossil insects in the diatoms of Shanwang. Bulletin of the Tianjin Institute of Geology and Mineral Resources 8:1-15 [D. Smith/E. Leckey/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 128550: authorized by Matthew Clapham, entered by Matthew Clapham on 11.06.2012, edited by Erin Leckey

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Arachnida
Araneae - Sparassidae	"Retina robusta" = Heteropoda robusta⁶ "Retina robusta" = Heteropoda robusta⁶ Hong 1985 spider 79104
Araneae - Tetragnathidae	"Eocryphoeca parva" = Tetragnatha parva⁶ "Eocryphoeca parva" = Tetragnatha parva⁶ Hong 1985 stretch spider 79101
Araneae - Araneidae	"Melanites orbiculatus" = Tethneus orbiculatus⁶, Araneus piceus⁶, Araneus ruidipedalis n. sp.⁶ "Melanites orbiculatus" = Tethneus orbiculatus⁶ Hong 1985 orb-weaving spider 79100 Araneus piceus⁶ Lin et al. 1989 orb-weaving spider 79099 (part of M. orbiculatus in Hong, 1985) Araneus ruidipedalis n. sp.⁶ Zhang et al. 1994 orb-weaving spider 79098
Insecta
Diptera - Tipulidae	Tipula insolita n. sp.⁵ Tipula insolita n. sp.⁵ Zhang 1989 crane fly 79027
Diptera - Bibionidae	Clothonopsis miocenica n. gen. n. sp.³, Plecia solaris n. sp.⁵, Plecia ludongensis n. sp.¹, "Bibio miocenica n. sp." = Bibio miocenicus¹, Bibio expansus⁵, Bibio cratopodus⁵, "Fungivora bullata n. sp." = Bibio bullatus⁵, Bibio brunneus n. sp.⁵, Bibio ventricosus n. sp.⁵, Bibio ruptineurus n. sp.⁵, Bibio deformatus n. sp.⁵, Bibio shanwangensis¹ Clothonopsis miocenica n. gen. n. sp.³ Hong and Wang 1987 love bug BNHM 80195 Plecia solaris n. sp.⁵ Zhang 1989 love bug 79125/6 (Bibio shanwangensis in Hong, 1985 - fig 26) Plecia ludongensis n. sp.¹ Hong and Wang 1985 love bug 80170 "Bibio miocenica n. sp." = Bibio miocenicus¹ Hong and Wang 1985 march fly 80117 Bibio expansus⁵ Zhang 1989 march fly 79053 (Bibio shanwangensis in Hong, 1985 - pl 7, fig 4) Bibio cratopodus⁵ Zhang 1989 march fly 79092 "Fungivora bullata n. sp." = Bibio bullatus⁵ Hong 1985 march fly 79003 Bibio brunneus n. sp.⁵ Zhang 1989 march fly 79120 (Bibio sp. in Hong, 1985) Bibio ventricosus n. sp.⁵ Zhang 1989 march fly 79033 (Bibio shanwangensis in Hong, 1985 - pl 7, figs 1, 2, 3) Bibio ruptineurus n. sp.⁵ Zhang 1989 march fly 79123 (Bibio sp. in Hong, 1985) Bibio deformatus n. sp.⁵ Zhang 1989 march fly 79057 (Bibio shanwangensis in Hong, 1985 - pl 4, fig 1) Bibio shanwangensis¹ Hong 1984 march fly 80123
Diptera - Sciaridae	Sciara brevipetiola n. sp.⁵ Sciara brevipetiola n. sp.⁵ Zhang 1989 dark-winged fungus gnat 79004
Odonata - Gomphaeschnidae	Jagoria sinica n. sp.⁵ Jagoria sinica n. sp.⁵ Zhang 1989 true dragonfly 79064 (Nasiaeschna orientalis in Hong, 1985)
Ephemeroptera - Heptageniidae	? Heptagenia sp.⁵ ? Heptagenia sp.⁵ Walsh 1863 stream mayfly 79095
Ephemeroptera - Leptophlebiidae	Paraleptophlebia sp.⁵ Paraleptophlebia sp.⁵ Lestage 1917 mayfly 79129 (Baetis sp. in Hong, 1985)
Dermaptera - Forficulidae	Apanechura asecta n. sp.⁵ Apanechura asecta n. sp.⁵ Zhang 1989 common earwig 79029
Hymenoptera - Cimbicidae	"Procimbex shanwangensis n. sp." = Abia shanwangensis, "Procimbex shandongense n. sp." = Abia shandongensis¹, Abia paurocephala n. sp.⁵ "Procimbex shanwangensis n. sp." = Abia shanwangensis Hong 1984 cimbicid sawfly 79038 "Procimbex shandongense n. sp." = Abia shandongensis¹ Hong and Wang 1985 cimbicid sawfly 80193 Abia paurocephala n. sp.⁵ Zhang 1989 cimbicid sawfly 79085
Hymenoptera - Xyelidae	"Megaxyela xiejiaheensis n. sp." = Xyelecia xiejiaheensis "Megaxyela xiejiaheensis n. sp." = Xyelecia xiejiaheensis Hong 1984 sawfly 79048
Hymenoptera - Ichneumonidae	Luia lacaris n. gen. n. sp.⁵ Luia lacaris n. gen. n. sp.⁵ Hong 1985 ichneumon wasp 79037
Hymenoptera - Vespidae	"Pseudoperla shanwangensis n. sp." = Eumenes shanwangensis "Pseudoperla shanwangensis n. sp." = Eumenes shanwangensis Hong 1984 potter wasp 79044
Hymenoptera - Tiphiidae	"Brachyopteris shanwangensis n. sp." = Tiphia shanwangensis "Brachyopteris shanwangensis n. sp." = Tiphia shanwangensis Hong 1984 wasp 79041
Hymenoptera - Thynnidae	Elis pyrula n. sp.⁵ Elis pyrula n. sp.⁵ Zhang 1989 wasp 79089
Hymenoptera - Formicidae	Paraphaenogaster shanwangensis n. sp., "Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster⁵, ? Tapinoma baculum n. sp.⁵, "Shanwangella palaeoptera n. gen. n. sp." = Liometopum palaeopterum⁵, Liometopum potamophilum n. sp.⁵, "Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis⁵, Lasius vetulus⁵, Lasius validus n. sp.⁵, ? Lasius truncatus n. sp.⁵, "Camponotus longus n. sp." = Camponotites longus⁵, "Camponotus shanwangense" = Camponotites shanwangensis, "Camponotus curviansatus n. sp." = Camponotites curviansatus⁵, "Camponotus microthoracus n. sp." = Camponotites microthoracus⁵, "Rabidia xiejiaheensis n. sp." = Camponotites xiejiaheensis, Camponotus mitis⁵, Formica linquensis n. sp.⁵ Paraphaenogaster shanwangensis n. sp. Hong 1984 ant 79047 "Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster⁵ Zhang 1989 ant 79054 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 3) ? Tapinoma baculum n. sp.⁵ Zhang 1989 ant 79051 "Shanwangella palaeoptera n. gen. n. sp." = Liometopum palaeopterum⁵ Zhang 1989 ant 79051 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 2) Liometopum potamophilum n. sp.⁵ Zhang 1989 ant 79036 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 6) "Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis⁵ Hong 1985 ant 79087; 80196 in Hong & Wang 1985 Lasius vetulus⁵ Dlussky 1981 ant 79030 Lasius validus n. sp.⁵ Zhang 1989 ant 79121 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 4) ? Lasius truncatus n. sp.⁵ Zhang 1989 ant 79055 "Camponotus longus n. sp." = Camponotites longus⁵ Zhang 1989 ant 79052 "Camponotus shanwangense" = Camponotites shanwangensis Hong 1984 ant 79031; 79043, 79076 (Liometopum jiaoyanshanense in Hong, 1985); ten specimens including 80179 in Hong & Wang 1985 "Camponotus curviansatus n. sp." = Camponotites curviansatus⁵ Zhang 1989 ant 79042 (Camponotus shanwangensis in Hong, 1985 - pl 27, fig 1) "Camponotus microthoracus n. sp." = Camponotites microthoracus⁵ Zhang 1989 ant 79046, 79127, 79128 "Rabidia xiejiaheensis n. sp." = Camponotites xiejiaheensis Hong 1984 ant 79069 Camponotus mitis⁵ Smith 1892 ant 79067 (Apis miocenica in Hing, 1985 - pl 22, fig 3) Formica linquensis n. sp.⁵ Zhang 1989 ant 79058 (Paraphaenogaster shanwangense in Hong, 1985 - pl 23, fig 4)
Hymenoptera - Apidae	Apis miocenica n. sp. Apis miocenica n. sp. Hong 1984 Chinese Miocene honey bee 79040, 79050; 80133 in Hong & Wang 1985
Hymenoptera - Pemphredonidae	"Sinostigma spinalata n. sp." = Pemphredon spinalatum "Sinostigma spinalata n. sp." = Pemphredon spinalatum Hong 1984 aphid wasp 79091
Coleoptera -	"Procarabus longipes n. sp." = Cistelites longipes⁵ "Procarabus longipes n. sp." = Cistelites longipes⁵ Hong 1985 beetle 79110
Coleoptera - Cupedidae	Cupes longus n. sp.³ Cupes longus n. sp.³ Hong and Wang 1987 reticulated beetle BNHM 80109
Coleoptera - Carabidae	"Shanwangicarabus furvus n. sp." = Chlaenius furvus², "Shanwangicarabus paucinus n. sp." = Chlaenius furvus², Scarites mancus n. sp.⁵, ? Carabus ovalus n. sp.², Calosoma cf. maderae⁵ "Shanwangicarabus furvus n. sp." = Chlaenius furvus² Hong 1984 ground beetle 80125, 80106, 80126, 480183 "Shanwangicarabus paucinus n. sp." = Chlaenius furvus² Hong 1984 ground beetle 80131 Scarites mancus n. sp.⁵ Zhang 1989 ground beetle 79106 (Aliscarabaeus granulatus in Hong, 1985) ? Carabus ovalus n. sp.² Hong and Wang 1986 ground beetle 80192 Calosoma cf. maderae⁵ Fabricius 1775 ground beetle 79001, 79002, 79108
Coleoptera -	Polyphaga indet.⁵ Polyphaga indet.⁵ Emery 1886 beetle 79063
Coleoptera - Staphylinidae	Lathrobium homorum n. sp.⁵ Lathrobium homorum n. sp.⁵ Zhang 1989 rove beetle 79021
Coleoptera -	Scarabaeoidea "indet. 1"⁵, Aliscarabaeus granulatus n. sp. Scarabaeoidea "indet. 1"⁵ Latreille 1802 scarab beetle 79045 (Magniscarabaeus furvus in Hong & Wang - pl 17, fig 2) Aliscarabaeus granulatus n. sp. Hong 1984 scarab beetle 79111
Coleoptera - Scarabaeidae	Anomala sp.⁵, Anomala synemosyna n. sp.⁵, Anomala lochmocola n. sp.⁵, "Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea⁵, "Magniscarabaeus furvus" = Anomala furva², Holotrichia spatha n. sp.⁵ Anomala sp.⁵ Samouelle 1819 scarab beetle 79096 Anomala synemosyna n. sp.⁵ Zhang 1989 scarab beetle 79006, 79007 Anomala lochmocola n. sp.⁵ Zhang 1989 scarab beetle 79112 "Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea⁵ Krell 2000 scarab beetle 79105; seven including 80101, 80120, 80172, 80188, 80184, 80194, mentioned in Hong & Wang 1987 "Magniscarabaeus furvus" = Anomala furva² Hong 1984 scarab beetle 80116, 80143 Holotrichia spatha n. sp.⁵ Zhang 1989 scarab beetle 79102
Coleoptera - Geotrupidae	Geotrupidae indet.⁵, "Obitiscarabaeus jiaoyanshanense n. gen. n. sp." = Geotrupes jiaoyanshanense Geotrupidae indet.⁵ Latreille 1802 earth-boring dung beetle 79107 (Magniscarabaeus brunneus in Hong, 1985 - pl 20, fig 3) "Obitiscarabaeus jiaoyanshanense n. gen. n. sp." = Geotrupes jiaoyanshanense Hong 1984 earth-boring dung beetle 79114/5
Coleoptera - Tenebrionidae	Sinocistela siphla n. gen. n. sp.⁵ Sinocistela siphla n. gen. n. sp.⁵ Zhang 1989 comb-clawed beetle 79028
Coleoptera - Meloidae	? Epicauta rufithoraca n. sp.⁵ ? Epicauta rufithoraca n. sp.⁵ Zhang 1989 blister beetle 79072
Coleoptera - Cerambycidae	Cerambycidae "indet. 1"⁵, Mesocacia pulla n. sp.⁵, Sinocalosoma longicornuta n. sp.², Mesosa soteria n. sp.⁵, Alifasciola daedalea n. sp.⁵ Cerambycidae "indet. 1"⁵ Latreille 1802 longhorned beetle 79020 Mesocacia pulla n. sp.⁵ Zhang 1989 longhorned beetle 79019 Sinocalosoma longicornuta n. sp.² Hong and Wang 1986 longhorned beetle 80175 Mesosa soteria n. sp.⁵ Zhang 1989 longhorned beetle 79008 Alifasciola daedalea n. sp.⁵ Zhang 1989 longhorned beetle 79074
Coleoptera - Curculionidae	Hylobius medianus n. sp.⁵, Hylobius aff. niitakensis⁵, Hylobius plenus⁵, "Sinocurculia granulata n. sp." = Atactogaster granulata, "Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis² Hylobius medianus n. sp.⁵ Zhang 1989 snout beetle 79117 Hylobius aff. niitakensis⁵ Kono 1933 snout beetle 79106, 79116 (ssp. fukienensis) Hylobius plenus⁵ Zhang 1989 snout beetle 79010, 79012, 79073 "Sinocurculia granulata n. sp." = Atactogaster granulata Hong 1984 snout beetle 79109 "Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis² Hong 1985 snout beetle 80128, 78001, 80107, 80134, 80173, 80174
Coleoptera - Brentidae	Longifistulia parva n. gen. n. sp.⁴ Longifistulia parva n. gen. n. sp.⁴ Hong and Wang 1987 weevil 80116
Coleoptera - Endomychidae	"Engonius alviolatus" = Gerstaeckerus alviolatus⁵ "Engonius alviolatus" = Gerstaeckerus alviolatus⁵ Hong 1984 handsome fungus beetle 79041
Coleoptera - Buprestidae	Agrilus corrugatus n. sp.⁵, Procarabus punctatus n. gen. n. sp. Agrilus corrugatus n. sp.⁵ Zhang 1989 jewel beetle 79062 Procarabus punctatus n. gen. n. sp. Hong 1984 jewel beetle 79118/9
Coleoptera - Dascillidae	Dascillus shandongianus⁵, Dascillus elongatus⁵ Dascillus shandongianus⁵ Zhang 1989 soft bodied plant beetle 79009, 79017, 79077, 79088 Dascillus elongatus⁵ Zhang 1989 soft bodied plant beetle 79081
Hemiptera - Aphididae	"Cinara reconditivenia n. sp." = Cinara (Cinara) reconditivenia⁵ "Cinara reconditivenia n. sp." = Cinara (Cinara) reconditivenia⁵ Zhang 1989 aphid 79056
Hemiptera -	Heteroptera indet.⁵ Heteroptera indet.⁵ Latreille 1810 true bug 79086
Hemiptera - Urostylididae	Urochela melaina n. sp.⁵, "Coreiopsis punctatus n. sp." = Urochela maculosa⁵ Urochela melaina n. sp.⁵ Zhang 1989 true bug 79024 "Coreiopsis punctatus n. sp." = Urochela maculosa⁵ Zhang 1989 true bug 79084
Hemiptera - Coreidae	"Coriopsis parva n. sp." = Coreiopsis parva⁴, Coreiopsis punctata n. gen. n. sp., "Coriopsis miocenica n. sp." = Coreiopsis miocenica⁴, Shanwangicoris longa n. gen. n. sp.⁴, Gonocerus shandongensis n. sp.⁵, Cloresmus shanwangi n. sp.⁵ "Coriopsis parva n. sp." = Coreiopsis parva⁴ Hong and Wang 1987 leaf-footed bug 80115 Coreiopsis punctata n. gen. n. sp. Hong 1984 leaf-footed bug 79084 "Coriopsis miocenica n. sp." = Coreiopsis miocenica⁴ Hong and Wang 1987 leaf-footed bug 80104 Shanwangicoris longa n. gen. n. sp.⁴ Hong and Wang 1987 leaf-footed bug 80129 Gonocerus shandongensis n. sp.⁵ Zhang 1989 leaf-footed bug 79025 Cloresmus shanwangi n. sp.⁵ Zhang 1989 leaf-footed bug 79005
Hemiptera - Belostomatidae	Manocerus stagnans n. gen. n. sp.⁵ Manocerus stagnans n. gen. n. sp.⁵ Zhang 1989 giant water bug 79016
Hemiptera - Nepidae	Coroides xiejiaheense n. gen. n. sp. Coroides xiejiaheense n. gen. n. sp. Hong 1984 true bug 79113
Hemiptera - Notonectidae	Tarsonecta mecopoda n. gen. n. sp.⁵ Tarsonecta mecopoda n. gen. n. sp.⁵ Zhang 1989 true bug 79013
Hemiptera - Cicadidae	"Oncotympana lapidescens n. sp." = Tanyocicada lapidescens⁵ "Oncotympana lapidescens n. sp." = Tanyocicada lapidescens⁵ Zhang 1989 cicada 79023
Hemiptera - Fulgoridae	Aphaena lithoecia n. sp.⁵, "Oxycephala xiejiaheensis" = Limois shanwangensis⁵, "Fulgoropsis fusca n. gen. n. sp." = Limois shanwangensis Aphaena lithoecia n. sp.⁵ Zhang 1989 fulgorid planthopper 79022 "Oxycephala xiejiaheensis" = Limois shanwangensis⁵ Hong 1979 fulgorid planthopper 79026, 79079, 79080 "Fulgoropsis fusca n. gen. n. sp." = Limois shanwangensis Hong 1979 fulgorid planthopper 79080

¹Hong and Wang 1985; ²Hong and Wang 1986; ³Hong and Wang 1987; ⁴Hong and Wang 1987; ⁵Zhang 1989; ⁶Zhang et al. 1994