El Breal de Orocual (Pliocene to of Venezuela)

Where: Monagas, Venezuela (9.8° N, 63.3° W: paleocoordinates 9.8° N, 63.0° W)

• coordinate stated in text

• local area-level geographic resolution

When: Mesa Formation, Late/Upper Pliocene to Late/Upper Pliocene (3.6 - 0.8 Ma)

• The Mesa Formation is early to middle Pleistocene age, based on thermoluminescence dates of 0.5 to 1.0 Ma (Carbón et al., 1992).

•Ruiz-Ramoni et al. 2017: El Breal de Orocual locality ORS-16 pit has been dated biochronologically to the Late Pliocene e Early Pleistocene based on the taxa Chapalmatherium sp., Pachyarmatherium leiseyi, and Smilodon gracilis (Rincon et al., 2009, 2011).

•Rincón et al. 2009: The deposit appears to be emplaced within the Mesa Formation, the age of which is assumed to be Pleistocene based on its stratigraphic position with respect to the Pliocene Las Piedras Formation (González de Juana, et al., 1980:713).

• group of beds-level stratigraphic resolution

Environment/lithology: terrestrial; tar and tar

• The faunal assemblage suggests that the El Breal de Orocual paleoenvironment was similar to that of the Venezuelan llanos today: an extensive savanna with rivers and patches of gallery forest (Rincón,et al., 2007, 2009).

•Ruiz-Ramoni et al. 2017: The fauna from ORS-16 consists of 25 mammal taxa including birds, testudines, and a crocodilian, as well as Cebus. All of these taxa indicate that the region was an extensive savanna with water bodies, and patches of gallery forest (Rincon et al., 2007, 2009, 2011; Fortier and Rincon, 2013). The presence of a capuchin monkey, very probably arboreal, supports this paleoecological reconstruction.

• This deposit has been interpreted as a tar seep deposit lying in the Mesa Formation (Hackley et al., 2006), a unit that consists of fluvio-deltaic strata.

•Rincón et al. 2009: The bones and sediments were clearly impregnated with asphalt at some time in the past, but no fluid asphalt is present in the deposit today. Solidified asphalt blocks are exposed in cross section in the wall of the trench in several places. These blocks are surrounded by a weathered corona of badly degraded wood, and appear to represent hollow cavities within logs or branches that were filled with asphalt that could not escape when the rest of the deposit was drained.

Size classes: macrofossils, mesofossils

Collection methods: Museum repositories: OR (El Breal de Orocual collection, hosted in Instituto Venezolano de Investigaciones Cient ́ıficas, Caracas, Venezuela)

Primary reference: A. D. Rincón, F. J. Prevosti, and G. E. Parra. 2011. New saber-toothed cat records (Felidae: Machairodontinae) for the Pleistocene of Venezuela, and the Great American Biotic Interchange. Journal of Vertebrate Paleontology 31(2):468-478 [C. Jaramillo/C. Suarez-Gomez]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 144977: authorized by Carlos Jaramillo, entered by Catalina Suarez-Gomez on 24.05.2013, edited by Philip Mannion and Grace Varnham

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Crocodylia -
Crocodylia indet.1 Owen 1842 crocodilian
Dorsal osteoderms, OR-1835, OR-1836; lateral osteoderm, OR- 343; centrum of cervical vertebra, OR-1833; centrum of first caudal vertebra, OR-1394; posterior caudal vertebra (OR-348); a set of uncatalogued fragments
 Crocodylia - Alligatoridae
Caimaninae indet.1, Caiman sp.1, "Caiman venezuelensis n. sp." = Caiman crocodilus1
Caimaninae indet.1 Brochu 1999 crocodilian
Posterior fragment of right jaw, OR-1679; anterior fragment of right jaw, OR-1683
Caiman sp.1 Spix 1825 caiman
Fragment of left dentary, OR-271; fragment of right maxilla, OR- 1678
"Caiman venezuelensis n. sp." = Caiman crocodilus1 Linnaeus 1758 Common caiman
OR-1677 - holotype - incomplete left premaxilla
Mammalia
 Didelphimorphia - Didelphidae
Didelphis sp.2 Linnaeus 1758 large American opossum
 Perissodactyla - Equidae
Hippidion sp.2 Owen 1869 horse
 Artiodactyla - Tayassuidae
Platygonus sp.2 Leconte 1848 peccary
Tayassu sp.2 Fischer 1814 peccary
 Carnivora - Canidae
cf. Protocyon indet.2 Giebel 1855 canine
 Carnivora - Felidae
Smilodon sp.2, Smilodon gracilis, "Homotheriini indet." = Homotheriina2, "Homotherium venezuelensis n. sp." = Xenosmilus venezuelensis
Smilodon sp.2 Lund 1841 saber-toothed cat
Smilodon gracilis Cope 1880 saber-toothed cat
Referred Materials: Upper incisor, OR-1346; fractured left upper crown of canine, 1342; left P3, 1344; right P3, 1347; right P4, 1343; left P4 fragments, 1345
"Homotheriini indet." = Homotheriina2 Jiangzuo et al. 2022 cat
"Homotherium venezuelensis n. sp." = Xenosmilus venezuelensis Rincón et al. 2011 cat
Holotype: OR-1352. Hypodigm: Three upper canines, OR-712, OR-1349, OR-1350; four upper canine fragments, OR-080, OR-258, OR-1219, OR-1386; right I3, OR-256; two right P4, OR- 255, OR-1390; two left dentaries, OR-259, OR-330; two dentary fragments, OR-710, OR-711; left lower canine, OR-254; two right lower canines, OR- 771, OR-1221; left m1, OR1389; left i3 and lower canine associate, OR-1166; right i1 crown, OR-081; left i1, OR-1385; right i2, OR-1381; right i3, OR-1220; left i3 crown, OR-785; two left p4, OR-257, OR-1384.
 Carnivora - Ursidae
cf. Arctotherium wingei2 Ameghino 1902 bear
 Primates - Cebidae
cf. Cebus sp.3 Erxleben 1777 capuchin monkey
 Rodentia -
Proechimys sp.2 Allen 1899 spiny rat
 Rodentia - Erethizontidae
Erethizontidae indet.2 Bonaparte 1845 porcupine
 Rodentia - Hydrochoeridae
"cf. Chapalmatherium indet." = Phugatherium2
"cf. Chapalmatherium indet." = Phugatherium2 Ameghino 1887 caviomorph
 Notoungulata - Toxodontidae
Mixotoxodon larensis2 Van Frank 1957 notoungulate
 Proboscidea -
Proboscidea indet.2 Illiger 1811 proboscidean
 Cingulata - Panochthidae
Hoplophorus sp.2 Lund 1838 edentate
 Cingulata - Pampatheriidae
Holmesina occidentalis2 Hoffstetter 1952 edentate
Pampatherium humboldtii2 Lund 1839 edentate
 Cingulata - Glyptodontidae
Glyptodon sp.2 Owen 1839 glyptodon
 Cingulata - Pachyarmatheriidae
cf. Pachyarmatherium leiseyi2 Downing and White 1995 edentate
 Cingulata - Dasypodidae
Propraopus sulcatus2 Lund 1838 armadillo
 Xenarthra - Myrmecophagidae
cf. Myrmecophaga sp.2 Linnaeus 1758 giant anteater
 Megatherioidea - Megatheriidae
Eremotherium sp.2 Spillmann 1948 edentate
 Megatherioidea - Megalonychidae
Megalonychidae indet.2 Gervais 1855 edentate