Insecta - Lepidoptera - Nymphalidae
Parent taxon: Papilionoidea according to H. Aiba et al. 2025
See also Carpenter 1992, Cockerell 1908, de Jong 2017, PeƱalver and Grimaldi 2006, Sohn et al. 2012 and van Nieukerken et al. 2011
Sister taxa: Archaeolycorea, Hesperiidae, Lithopsyche, Lycaenidae, Papilionidae, Pieridae, Riodinidae, Stolopsyche
Subtaxa: Apatura Biblidinae Chlorippe Danainae Eugonia Libytheinae Limenitidinae Nymphalinae Nymphalis Satyrinae
Ecology:
Distribution:
• Quaternary of Japan (1 collection)
• Pliocene to Pleistocene of Japan (1)
• Pliocene of Japan (1)
• Miocene of Croatia (2), the Dominican Republic (1), Germany (2), the Russian Federation (1)
• Oligocene of Brazil (1), France (5), United States (1: Montana)
• Eocene of the Russian Federation (1), the United Kingdom (1), United States (8: Colorado)
Total: 26 collections including 32 occurrences