Family Nymphalidae Rafinesque 1815 (brush-footed butterfly)

Insecta - Lepidoptera - Nymphalidae

Parent taxon: Papilionoidea according to H. Aiba et al. 2025

See also Carpenter 1992, Cockerell 1908, de Jong 2017, PeƱalver and Grimaldi 2006, Sohn et al. 2012 and van Nieukerken et al. 2011

Sister taxa: Archaeolycorea, Hesperiidae, Lithopsyche, Lycaenidae, Papilionidae, Pieridae, Riodinidae, Stolopsyche

Subtaxa: Apatura Biblidinae Chlorippe Danainae Eugonia Libytheinae Limenitidinae Nymphalinae Nymphalis Satyrinae

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Ecology:

Distribution:

• Quaternary of Japan (1 collection)

• Pliocene to Pleistocene of Japan (1)

• Pliocene of Japan (1)

• Miocene of Croatia (2), the Dominican Republic (1), Germany (2), the Russian Federation (1)

• Oligocene of Brazil (1), France (5), United States (1: Montana)

• Eocene of the Russian Federation (1), the United Kingdom (1), United States (8: Colorado)

Total: 26 collections including 32 occurrences

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