Full reference: A. Handlirsch. 1906. Die Fossilen Insekten und die Phylogenie der Rezenten Formen, parts I-IV. Ein Handbuch fur Palaontologen und Zoologen 1-640
Parent taxon: Scarabaeoidea according to G. V. Nikolajev 2008
See also Carpenter 1992, Hong 1992, Hong and Wang 1990, Krell 2000, Nikolajev 1992 and Ponomarenko 1986
Sister taxa: Aliscarabaeus, Alloioscarabaeidae, Ateuchites, Belohinidae, Bolboceratidae, Ceafornotensis, Copris subterraneus, Cretaclopus, Cretonitis, Diphyllostomatidae, Geotrupidae, Glaphyridae, Glaresidae, Gymnopleurus deperditus, Holcorobeini, Holcorobeus, Hongscarabaeus, Hybosoridae, Hybosorites, Juraclopus, Lucanidae, Mahengea, Mesaclopus, Mesoscarabaeus, Mongolrobeus, Ochodaeidae, Onthophagus urusheeri, Opiselleipon, Passalidae, Passalopalpidae, Pleocomidae, Prionocephale, Prophaenognatha, Scarabaeidae, Scelocopris, Septiventeridae, Trogidae
Subtaxa: Geotrupoides jiaoheense Geotrupoides kezuoensis Geotrupoides lithographicus Geotrupoides saxosus Geotrupoides songyingziensis
Ecology:
Distribution:
• Cretaceous of China (4 collections)
• Jurassic of Germany (1)
Total: 5 collections each including a single occurrence
Specimen images are retrieved through the ePANDDA API.
Click image to enlarge. Click to access iDigBio record.