Insecta - Megasecoptera - Scytohymenidae
Full reference: A. V. Martynov. 1937. Permian fossil insects from Kargala and their relationships. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7:1-92
Parent taxon: Megasecoptera according to M. Pecharová and J. Prokop 2018
See also Carpenter 1992, Martynov 1937 and Shcherbakov 2009
Sister taxa: Adiaphtharsia, Alectoneuridae, Anchineuridae, Ancopteridae, Archaeoscolex, Archaeoscolex corneus, Arcioneuridae, Aspidohymenidae, Aspidothoracidae, Bardohymenidae, Brodiidae, Caulopteridae, Corydaloididae, Dictyoneurellidae, Engisopteridae, Eumegasecoptera, Foririidae, Ischnoptilidae, Lameereites, Mischopteridae, Moravohymenidae, Piesbergpteridae, Protohymenidae, Sphecopteridae, Sphecorydaloididae, Vorkutiidae, Xenopteraidae, Raphidiopsis
Subtaxa: Karoohymen Oceanoptera Scytohymen Tshekardohymen
Type: Scytohymen
Ecology:
Distribution:
• Permian of the Russian Federation (4 collections), South Africa (1)
Total: 5 collections including 7 occurrences