Tribe Gymnopleurini Lacordaire 1856 (scarab beetle)
Insecta - Coleoptera - Scarabaeidae
Parent taxon: Scarabaeinae according to P. Bouchard et al. 2011
Sister taxa: Anachalcos, Ateuchini, Canthochilum, Coprini, Deltochilini, Heliocopris, Metacatharsius, Oniticellini, Onitini, Onthophagini, Onthophagus spitsbergeniensis, Phanaeini, Scarabaeini
Subtaxa: Gymnopleurus
Ecology:
Distribution: found only at Oeningen (Zurich collection) (Miocene of Germany)