Insecta - Coleoptera - Scarabaeidae
Full reference: P. A. Latreille. 1802. Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes. 3:1-467
Parent taxon: Scarabaeidae according to F. Tello et al. 2023
See also Bouchard et al. 2011, Cai et al. 2022, Krell 2000, Lawrence and Newton 1995, Nikolajev 2007, Tarasov and Dimitrov 2016, Tarasov et al. 2016 and Wappler 2003
Sister taxa: none
Subtaxa: Anachalcos Ateuchini Canthochilum Coprini Deltochilini Gymnopleurini Heliocopris Metacatharsius Oniticellini Onitini Onthophagini Onthophagus spitsbergeniensis Phanaeini Scarabaeini
Type: Scarabaeus
Ecology:
Distribution:
• Quaternary of Chile (2 collections), Ecuador (1), Japan (7), Peru (1), the United Kingdom (9), United States (6: California, Pennsylvania, Virginia)
• Pliocene of Georgia (1), Germany (1), Japan (3)
• Miocene of China (1), the Dominican Republic (1), Germany (2), Japan (1), Kenya (2)
• Oligocene of France (1), Germany (1)
• Eocene of France (1), Germany (2), the United Kingdom (1)
• Paleocene of Svalbard and Jan Mayen (1)
Total: 45 collections including 69 occurrences