Insecta - Coleoptera - Scarabaeidae

Parent taxon: Scarabaeinae according to P. Bouchard et al. 2011

Sister taxa: Anachalcos, Ateuchini, Canthochilum, Coprini, Deltochilini, Gymnopleurini, Heliocopris, Metacatharsius, Onitini, Onthophagini, Onthophagus spitsbergeniensis, Phanaeini, Scarabaeini

Subtaxa: Drepanocerina Oniticellina

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Ecology:

Distribution: found only at Trafalgar Square (Quaternary of the United Kingdom)

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