Actinopteri - Cypriniformes - Cyprinidae
Parent taxon: Cyprinidae according to J. S. Nelson et al. 2016
See also Böhme 2010 and Hay 1902
Sister taxa: Acheilognathinae, Acrocheilus, Alisodon, Amblypharyngodon, Amyzon, Aspius, Barbinae, Barbus, Cultrinae, Cyprinion, Cyprinus, Danioninae, Ecocarpia, Esox destructus, Gila, Gobio, Gobioninae, Hexapsephus, Iquius, Labeoninae, Lavinia, Leuciscinae, Leuciscus, Mylocyprinus, Nemachilus, Opsariichthyinae, Orthodon, Palaeoleuciscus, Palaeotinca, Phoxininae, Planktophaga, Protothymallus, Ptychocheilus, Puntius, Rhodeus, Siphateles, Squaliobarbinae, Thynichthys, Tinca, Tincinae, Xenocyprinae
Subtaxa: Aphelichthys Diastichus Notropis Palaeocarassius Rutilus
Ecology: nektonic omnivore
Distribution:
• Quaternary of the Russian Federation (1 collection), the United Kingdom (1), United States (1: South Dakota)
• Pliocene of Germany (1), Moldova (1), the Russian Federation (1)
• Miocene of Germany (5), Moldova (1), the Russian Federation (1), Spain (2), Switzerland (2), Ukraine (1), United States (1: Kansas)
Total: 19 collections including 20 occurrences