Actinopteri - Cypriniformes - Cyprinidae

Parent taxon: Cyprinidae according to J. S. Nelson et al. 2016

See also Vasilyan and Carnevale 2013

Sister taxa: Acheilognathinae, Acrocheilus, Alisodon, Amblypharyngodon, Amyzon, Aspius, Barbinae, Barbus, Cultrinae, Cyprininae, Cyprinion, Cyprinus, Danioninae, Ecocarpia, Esox destructus, Gila, Gobio, Gobioninae, Hexapsephus, Idadon, Iquius, Lavinia, Leuciscinae, Leuciscus, Mylocyprinus, Nemachilus, Opsariichthyinae, Orthodon, Palaeoleuciscus, Palaeotinca, Phoxininae, Planktophaga, Protothymallus, Ptychocheilus, Puntius, Rhodeus, Siphateles, Squaliobarbinae, Thynichthys, Tinca, Tincinae, Xenocyprinae

Subtaxa: Garraini

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Ecology: nektonic omnivore

Distribution: found only at Jradzor (Pliocene of Armenia)

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