Insecta - Coleoptera - Carabidae
Alternative spelling: Steropus
Synonyms: Pterostichus (Steroderus) Motschulsky 1850, Steroderus Motschulsky 1850
Full reference: P. F. M. Dejean. 1821. Catalogue de la collection de Coléoptères de M. le Baron Dejean 1-136
Parent taxon: Pterostichus according to A. M. Lomnicki 1894
See also Clapham 2023 and Dejean 1821
Sister taxa: Pterostichus (Argutor), Pterostichus (Bothriopterus), Pterostichus (Cryobius), Pterostichus (Eosteropus), Pterostichus (Hypherpes), Pterostichus (Lamenius), Pterostichus (Lenapterus), Pterostichus (Leptoferonia), Pterostichus (Melanius), Pterostichus (Metallophilus), Pterostichus (Phonias), Pterostichus (Pseudomaseus), Pterostichus abnormis, Pterostichus abrogatus, Pterostichus cancellatus, Pterostichus costatus, Pterostichus depletus, Pterostichus destitutus, Pterostichus destructus, Pterostichus distinctus, Pterostichus dormitans, Pterostichus fernquisti, Pterostichus fractus, Pterostichus gelidus, Pterostichus gracilis, Pterostichus gurnetensis, Pterostichus laevigatus, Pterostichus melanarius, Pterostichus minor, Pterostichus minutulus, Pterostichus molestus, Pterostichus niger, Pterostichus nigrita, Pterostichus parens, Pterostichus pumpellyi, Pterostichus strenuus, Pterostichus stygicus, Pterostichus sublaevis, Pterostichus tundrae, Pterostichus vernalis, Pterostichus vetustus, Pterostichus walcotti
Subtaxa: Pterostichus (Steropus) aethiops
Ecology:
Distribution: found only at Boryslawia (Quaternary of Ukraine)