Insecta - Coleoptera - Carabidae
Parent taxon: Pterostichus according to Y. Bousquet 2012
Sister taxa: Pterostichus (Argutor), Pterostichus (Bothriopterus), Pterostichus (Cryobius), Pterostichus (Eosteropus), Pterostichus (Hypherpes), Pterostichus (Lamenius), Pterostichus (Leptoferonia), Pterostichus (Melanius), Pterostichus (Metallophilus), Pterostichus (Phonias), Pterostichus (Pseudomaseus), Pterostichus (Steropus), Pterostichus abnormis, Pterostichus abrogatus, Pterostichus cancellatus, Pterostichus costatus, Pterostichus depletus, Pterostichus destitutus, Pterostichus destructus, Pterostichus distinctus, Pterostichus dormitans, Pterostichus fernquisti, Pterostichus fractus, Pterostichus gelidus, Pterostichus gracilis, Pterostichus gurnetensis, Pterostichus laevigatus, Pterostichus melanarius, Pterostichus minor, Pterostichus minutulus, Pterostichus molestus, Pterostichus niger, Pterostichus nigrita, Pterostichus parens, Pterostichus pumpellyi, Pterostichus strenuus, Pterostichus stygicus, Pterostichus sublaevis, Pterostichus tundrae, Pterostichus vernalis, Pterostichus vetustus, Pterostichus walcotti
Subtaxa: Pterostichus (Lenapterus) agonus Pterostichus (Lenapterus) punctatissimus Pterostichus (Lenapterus) vermiculosus
Ecology:
Distribution:
• Quaternary of Greenland (5 collections), the Russian Federation (5), United States (1: Pennsylvania)
• Miocene of United States (1: Alaska)
Total: 12 collections including 13 occurrences