Aves - Suliformes - Sulidae
Parent taxon: Suliformes according to J. F. Clements et al. 2017
See also Bourdon et al. 2005, Brodkorb 1955, Carroll 1988, Case 1904, Chandler 1990, Cheneval 1984, Cheneval 1993, del Hoyo et al. 2014, Figueiredo et al. 2023, Gohlich 2003, Hay 1902, Hay 1930, Howard 1968, Howard 1972, Howard 1978, Howard 1982, Howard and Barnes 1987, Lambrecht 1933, Livezey and Zusi 2007, Mayr 2002, Mayr 2005, Mayr 2005, Mayr 2011, Mayr 2022, Miller and Bowman 1958, Mlikovsky 2007, Olson 1975, Olson 1985, Olson 1985, Olson and Hasegawa 1996, Olson and James 1982, Olson and Rasmussen 2001, Steadman et al. 1988, Stucchi 2003, Stucchi and Urbina 2004, Wetmore 1940, Wetmore 1956 and Zittel 1890
Sister taxa: Anhingidae, Fregatidae, Mangystania, Phalacrocoracidae, Plotopteridae
Subtaxa: Empheresula Enkurosula Eostega Masillastega Microsula Miosula Morus Paleosula Papasula Prophalacrocorax Ramphastosula Sarmatosula Sula
Type: Sula
Ecology: ground dwelling carnivore
Distribution:
• Quaternary of American Samoa (1 collection), Australia (2), the Bahamas (1), Brazil (1), Cook Islands (1), French Polynesia (4), Greece (1), Italy (1), the Marshall Islands (2), Mauritius (2), Micronesia (2), New Zealand (1), Norfolk Island (9), Pitcairn (5), Saint Helena (13), Solomon Islands (22), South Africa (3), Spain (3), Ukraine (1), the United Kingdom (10), United States (11: California, Florida, Virginia), Vanuatu (1), Venezuela (6)
• Blancan of United States (7: California)
• Pliocene of Morocco (1), Peru (1), South Africa (1), United States (5: California, North Carolina)
• Miocene of Austria (1), Chile (1), France (2), Germany (1), Japan (1), Mexico (5), Peru (9), Portugal (1), Romania (1), United States (28: California, Delaware, Florida, Maryland, New Jersey, North Carolina, Virginia)
• Oligocene of France (2), Germany (1)
• Eocene of Germany (1), Romania (1), the United Kingdom (1)
Total: 174 collections including 244 occurrences