Family Passalidae Leach 1815 (bess beetle)

Insecta - Coleoptera - Passalidae

Alternative spelling: Passalida

Full reference: W. E. Leach. 1815. Entomology. The Edinburgh Encyclopaedia 9:57-172

Parent taxon: Scarabaeoidea according to P. Bouchard et al. 2011

See also Carpenter 1992, Cockerell 1927, Krell 2000 and Lawrence and Newton 1995

Sister taxa: Acanthoceridae, Aliscarabaeus, Alloioscarabaeidae, Ateuchites, Belohinidae, Copris subterraneus, Cretonitis, Diphyllostomatidae, Eremazidae, Geotrupidae, Glaphyridae, Glaresidae, Gymnopleurus deperditus, Hongscarabaeus, Hybosoridae, Hybosorites, Lithoscarabaeidae, Lucanidae, Mesoscarabaeus, Ochodaeidae, Onthophagus urusheeri, Opiselleipon, Paralucanidae, Passalopalpidae, Pleocomidae, Prionocephale, Scarabaeidae, Scelocopris, Septiventeridae, Trogidae

Subtaxa: Aulacocyclinae Passalinae

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Type: Passalus

Distribution:

• Miocene of China (1 collection)

• Oligocene of United States (1: Oregon)

• Cretaceous of Myanmar (2)

Total: 4 collections including 5 occurrences

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