Family Passalidae Leach 1815 (bess beetle)

Insecta - Coleoptera - Passalidae

Alternative spelling: Passalida

Full reference: W. E. Leach. 1815. Entomology. The Edinburgh Encyclopaedia 9:57-172

Parent taxon: Scarabaeoidea according to C. Y. Cai et al. 2022

See also Bouchard et al. 2011, Carpenter 1992, Cockerell 1927, Krell 2000 and Lawrence and Newton 1995

Sister taxa: Aliscarabaeus, Alloioscarabaeidae, Ateuchites, Belohinidae, Bolboceratidae, Ceafornotensis, Copris subterraneus, Crassiungula, Cretaclopus, Cretonitis, Diphyllostomatidae, Geotrupidae, Geotrupoides, Glaphyridae, Glaresidae, Gymnopleurus deperditus, Holcorobeini, Holcorobeus, Hongscarabaeus, Hybosoridae, Hybosorites, Juraclopus, Lucanidae, Mahengea, Mesaclopus, Mesoscarabaeus, Mongolrobeus, Ochodaeidae, Onthophagus urusheeri, Opiselleipon, Passalopalpidae, Pleocomidae, Prionocephale, Prophaenognatha, Scarabaeidae, Scelocopris, Septiventeridae, Trogidae

Subtaxa: Aulacocyclinae Passalinae

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Type: Passalus

Distribution:

• Miocene of China (1 collection)

• Oligocene of United States (1: Oregon)

• Cretaceous of Brazil (1), Myanmar (2)

Total: 5 collections including 6 occurrences

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