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Testudo nanus

Osteichthyes - Testudines - Testudinidae

Taxonomy
Testudo nanus was named by Gilmore (1931). Its type specimen is AMNH 6692, a dermal scale, and it is a 3D body fossil. Its type locality is Twin Obo (Ulan Gochu horizon), which is in an Oligocene terrestrial horizon in the Ulan Gochu Formation of China.

It was recombined as Indotestudo nanus by Crumly (1983).

Sister species lacking formal opinion data
T. antiqua, T. aralensis, T. chailloti, T. csakvarensis, T. lambrechti, T. lemanensis, T. marmoreum, T. mellingi, T. noviciensis, T. opisthoklitea, T. peltastes, T. pyrenaica, T. rectogularis, T. risgoviensis, T. roquesi, T. turgaica, T. pardalis
Entered
by M. Carrano on 2010-11-11; modified by D. Nicholson on 2016-05-19

Synonymy list
YearName and author
1931Testudo nanus Gilmore p. 241 figs. 21-22; Pl. 7
1963Testudo nanus Yeh
1983Indotestudo nanus Crumly
2008Testudo nanus Brinkman et al. p. 93

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
RankNameAuthor
classReptilia
subclassEureptilia()
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—ArchelosauriaCrawford 2015
—Pantestudines
—Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
—Pantestudinoidea
superfamilyTestudinoidea
familyTestudinidaeBatsch 1788
subfamilyTestudininae
tribeTestudininiBatsch 1788
genusTestudo
speciesnanus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Testudo nanus Gilmore 1931
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Diagnosis
ReferenceDiagnosis
C. W. Gilmore 1931Small size; shell depremsed, quadrangular in outline; surfaces smooth other than costals being crossed by faint longitudinal grooves and ridges; anterior margin shallowly excavated; sulci shallowly impressed; marginal sulcus, coinciding with costo-peripheral suture except perhaps in front where it appears to run below; supracaudal scute divided; front lobe of plastron strongly produced; pecto-humeral sulcus crossing on anterior half of entoplastron; entoplastron sub-ovate, and largely behind the base line of the anterior lobe; gular scutes in front of entoplastron; pectoral scute subequal in length to abdominal; posterior lobe shallowly notched; lobe composed entirely of xiphiplastral bones; movable.