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Chimaeriformes (chimaera)
Taxonomy
Chimaeriformes was named by Obruchev (1953). It is extant.
It was assigned to Holocephali by Applegate (1970), Duffin and Furrer (1981), Duffin (1981), Purdy et al. (2001); to Chondrichthyes by Sepkoski (2002); to Holocephali by Kriwet and Gazdizicki (2003); to Holocephali by Thurmond and Jones (1981), Cvancara and Hoganson (1993), Delsate et al. (2002), Kent (2018); to Chimaerimorpha by van der Laan (2019); to Holocephalomorpha by Long (2011), Gouiric-Cavalli et al. (2015), Nelson et al. (2016), Popov et al. (2020); and to Holocephalimorpha by Nelson (2006), Ebersole et al. (2022), Popov et al. (2025).
It was assigned to Holocephali by Applegate (1970), Duffin and Furrer (1981), Duffin (1981), Purdy et al. (2001); to Chondrichthyes by Sepkoski (2002); to Holocephali by Kriwet and Gazdizicki (2003); to Holocephali by Thurmond and Jones (1981), Cvancara and Hoganson (1993), Delsate et al. (2002), Kent (2018); to Chimaerimorpha by van der Laan (2019); to Holocephalomorpha by Long (2011), Gouiric-Cavalli et al. (2015), Nelson et al. (2016), Popov et al. (2020); and to Holocephalimorpha by Nelson (2006), Ebersole et al. (2022), Popov et al. (2025).
Synonymy list
| Year | Name and author |
|---|---|
| 1953 | Chimaeriformes Obruchev |
| 1970 | Chimaeriformes Applegate p. 389 |
| 1981 | Chimaeriformes Duffin p. 470 |
| 1981 | Chimaeriformes Duffin and Furrer p. 810 |
| 1981 | Chimaeriformes Thurmond and Jones p. 77 |
| 1993 | Chimaeriformes Cvancara and Hoganson p. 12 |
| 2001 | Chimaeriformes Purdy et al. p. 161 |
| 2002 | Chimaeriformes Delsate et al. p. 25 |
| 2002 | Chimaeriformes Sepkoski |
| 2003 | Chimaeriformes Kriwet and Gazdizicki p. 34 |
| 2006 | Chimaeriformes Nelson p. 45 |
| 2011 | Chimaeriformes Long p. 242 |
| 2015 | Chimaeriformes Gouiric-Cavalli et al. p. 4 |
| 2016 | Chimaeriformes Nelson et al. p. 51 |
| 2018 | Chimaeriformes Kent p. 54 |
| 2019 | Chimaeriformes van der Laan p. 39 |
| 2020 | Chimaeriformes Popov et al. p. 5 |
| 2022 | Chimaeriformes Ebersole et al. p. 373 |
| 2025 | Chimaeriformes Popov et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Or. Chimaeriformes Obruchev 1953 [chimaera]
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Fm. Callorhynchidae Garman 1901 [plownose chimaera]
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Subfm. Callorhynchinae Garman 1901
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G. †Ptyktoptychion Lees 1986
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†Ptyktoptychion tayyo Lees 1986
†Ptyktoptychion wadeae Bartholomai 2008
G. †Elasmodus Egerton
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†Elasmodus avirostrus Averianov 1999
†Elasmodus greenoughi Agassiz 1838
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Invalid names: Aptychus crassus Hebert 1855 [synonym], Elasmodus crassus Leriche 1906 [synonym], Elasmodus planus Leriche 1929 [synonym]
†Elasmodus kempi Ward 1977
†Elasmodus khosatzkyi Averianov 1999
†Elasmodus rossicus Averianov 1999
†Elasmodus sinzovi Averianov and Glickman 1994
†Elasmodus tsheganicus Averianov 2001
†Elasmodus zharyk Averianov 1999
Invalid names: Psaliodus Agassiz 1843 [synonym]
G. †Ischyodus Egerton 1843
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†Ischyodus avitus Meyer 1800
†Ischyodus beaumonti Egerton 1843
†Ischyodus bifurcatus Case 1978
†Ischyodus bohemicus Bayer 1902
†Ischyodus crassus Woodward and Withe 1930
†Ischyodus dolloi Leriche 1902
†Ischyodus dufrenoyi Egerton 1843
†Ischyodus egertoni Buckland 1835
†Ischyodus emarginatus Egerton 1843
†Ischyodus incisus Newton 1878
†Ischyodus latus Newton 1878
†Ischyodus mortoni Chapman and Pritchard 1907
†Ischyodus panus Newton 1878
†Ischyodus quenstedti Wagner 1857
†Ischyodus rayhaasi Hoganson and Erickson 2005
†Ischyodus thurmanni Pictet and Campiche 1878
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Invalid names: Ischyodus brevirostris Newton 1876 [synonym]
†Ischyodus townsendi Buckland 1835
†Ischyodus yanschini Averianov 1991
†Ischyodus zinsmeisteri Applegate 1975
Subor. Chimaeroidei
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G. †Anteliochimaera Tanaka 1909
Fm. Callorhinchidae Garman 1901 [plownose chimaera]
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G. Callorhinchus Lacépède 1798
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†Callorhinchus alfordi Cicimurri and Ebersole 2015
†Callorhinchus phillipsi Cicimurri and Ebersole 2015
Callorhinchus stahli Kriwet and Gazdizicki 2003
G. †Ganodus Agassiz 1843
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†Chimaera (Ganodus) colei Buckland 1835
†Chimaera (Ganodus) curvidens Agassiz 1843
†Chimaera (Ganodus) neglecta Agassiz 1843
†Chimaera (Ganodus) owenii Buckland 1835
†Chimaera (Ganodus) rugulosa Agassiz 1843
†Ganodus dentatus Egerton 1847
†Ganodus oweni Agassiz 1843
†Ganodus rugulosus Egerton 1847
G. †Pachymylus Woodward 1892
Superfm. Callorhinchoidea
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G. †Chalcodus Zittel 1887
Fm. Chimaeridae
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G. Chimaera Linnaeus 1758
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Subg. †Chimaera (Ischyodon) Agassiz 1843
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†Chimaera (Ischyodon) beaumontii Agassiz 1843
†Chimaera (Ischyodon) brevirostris Agassiz 1843
†Chimaera (Ischyodon) dufrenoyi Agassiz 1843
†Chimaera (Ischyodon) dutertrii Agassiz 1843
†Chimaera (Ischyodon) emarginata Agassiz 1843
†Chimaera (Ischyodon) helvetica Agassiz 1843
†Chimaera (Ischyodon) johnsonii Agassiz 1843
†Chimaera (Ischyodon) tessoni Buckland 1835
†Ischyodon bucklandi Agassiz 1843
Subg. †Chimaera (Psittacodon) Agassiz 1843
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†Chimaera (Psittacodon) falcata Agassiz 1843
†Chimaera (Psittacodon) psttacina Agassiz 1843
†Chimaera furcata Fritsch 1878
†Chimaera pliocenica Woodward 1891
†Chimaera seymourensis Ward and Grande 1991
†Chimaera venablesae Casier 1966
G. †Hedronchus Cope 1876
G. †Hydrolagus Gill 1862
G. †Leptomylus Cope 1869
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†Leptomylus cookii Cope 1870
†Leptomylus densus Cope 1869
†Leptomylus forfex Cope 1875
Superfm. Chimaeroidea
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G. †Chimaerotheca Brown 1946
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†Chimaerotheca alaskana Brown 1946
†Chimaerotheca besselsi Brown 1946
†Chimaerotheca germanica Brown 1946
†Chimaerotheca montanana Brown 1946
†Chimaerotheca newmexicana Brown 1946
†Chimaerotheca reperere Gottfried and Fordyce 2015
†Chimaerotheca wyomingana Brown 1946
Fm. †Edaphodontidae Owen 1846
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G. †Edaphodon Buckland 1838
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†Edaphodon agassizi Buckland 1835
†Edaphodon antwerpiensis Leriche 1926
†Edaphodon barberi Applegate 1970
†Edaphodon crassus Newton 1878
†Edaphodon divaricatus Cope 1869
†Edaphodon eocaenus Woodward 1891
†Edaphodon fecundus Cope 1875
†Edaphodon hesperis Shin 2010
†Edaphodon incrassatus Cope 1875
†Edaphodon kawai Consoli 2006
†Edaphodon laminosus Newton 1878
†Edaphodon laterigerus Cope 1869
†Edaphodon latidens Cope 1875
†Edaphodon leptognathus Agassiz 1843
†Edaphodon mantellii Buckland 1835
†Edaphodon miersii Marsh 1869
†Edaphodon mirabilis Chapman and Cudmore 1924
†Edaphodon mirificus Leidy 1856
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Invalid names: Ischyodus gaskillii Cope 1875 [synonym], Ischyodus monolophus Cope 1869 [synonym]
†Edaphodon reedi Newton 1878
†Edaphodon sedgwicki Agassiz 1838
†Edaphodon smockii Cope 1869
†Edaphodon snowhillensis Gouiric-Cavalli et al. 2015
†Edaphodon solidulus Cope 1869
†Edaphodon stenobryus Cope 1875
†Edaphodon sweeti Chapman and Pritchard 1907
Invalid names: Diphrissa Cope 1875 [synonym], Dipristis Marsh [synonym]
G. †Icthypriapus
Subor. †Myriacanthoidei Patterson 1965
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Fm. †Chimaeropsidae
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Fm. †Myriacanthidae Woodward 1889
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G. †Halonodon Duffin 1984
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†Halonodon luxembourgensis Duffin and Delsate 1993
†Halonodon warneri Duffin 1984
G. †Myriacanthus Agassiz 1836
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†Myriacanthus retrorsus Agassiz 1837
Invalid names: Prognathodus Egerton 1872 [synonym]
Invalid names: Acanthorhinidae Patterson 1965 [synonym]
Invalid names: Chimeroidei Patterson 1965 [empty], Myracanthiodei [empty], Squalorajoidei Patterson 1965 [empty]
Diagnosis
No diagnoses are available