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Geochelonini
Discussion
Named first by C. R. Crumly in his 1984 thesis
Taxonomy
Geochelonini was named by Swingland and Klemens (1989). It is extant.
It was assigned to Geocheloninae by Swingland and Klemens (1989); to Testudininae by Vlachos (2021); and to Testudinidae by Vlachos et al. (2022), Pappa et al. (2023).
It was assigned to Geocheloninae by Swingland and Klemens (1989); to Testudininae by Vlachos (2021); and to Testudinidae by Vlachos et al. (2022), Pappa et al. (2023).
Subtaxa
Synonymy list
| Year | Name and author |
|---|---|
| 1989 | Geochelonini Swingland and Klemens p. 124 |
| 2021 | Geochelonini Vlachos |
| 2022 | Geochelonini Vlachos et al. |
| 2023 | Geochelonini Pappa et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Tr. Geochelonini Swingland and Klemens 1989
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G. Chelonoidis Fitzinger 1835
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†Chelonoidis abingdonii Gunther 1877 [Pinta giant tortoise]
†Chelonoidis alburyorum Franz and Franz 2009
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†Chelonoidis alburyorum alburyorum Franz and Franz 2009
†Chelonoidis alburyorum keegani Franz et al. 2020
†Chelonoidis alburyorum sementis Franz et al. 2020
Chelonoidis becki Rothschild 1901 [Volcan wolf giant tortoise]
Chelonoidis carbonarius Spix 1824 [Red-footed tortoise]
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Invalid names: Geochelone atlantica López-Jurado et al. 1998 [synonym]
Chelonoidis chathamensis Van Denburgh 1907 [San Cristobal giant tortoise]
Chelonoidis chilensis Gray 1870 [Chaco tortoise]
†Chelonoidis cubensis Leidy 1868
Chelonoidis darwini Van Denburgh 1907 [Santiago giant tortoise]
Chelonoidis denticulata Linnaeus 1766 [Yellow-footed tortoise]
†Chelonoidis dominicensis Albury et al. 2018
Chelonoidis donosobarrosi Freiberg 1973
Chelonoidis duncanensis Garman 1996 [Pinzon giant tortoise]
†Chelonoidis gallardoi Rovereto 1914
†Chelonoidis gersoni Viñola-López and Almonte 2022
†Chelonoidis gringorum Simpson 1942
†Chelonoidis hesterna Auffenberg 1971
Chelonoidis hoodensis Van Denburgh 1907 [Espanola giant tortoise]
†Chelonoidis lutzae Zacarias et al. 2013
†Chelonoidis marcanoi Turvey et al. 2017
†Chelonoidis meridiana Vlachos et al. 2022
†Chelonoidis monensis Williams 1952
Chelonoidis nigra Quoy and Gaimard 1824 [Floreana giant tortoise]
Chelonoidis petersi Freiberg 1973
†Chelonoidis petrocellii Agnolin 2021
†Chelonoidis phantastica Van Denburgh 1907 [Fernandina giant tortoise]
Chelonoidis porteri Rothschild 1903 [Santa Cruz giant tortoise]
Chelonoidis vicina Gunther 1875 [Southern Isabela giant tortoise]
G. †Titanochelon Pérez-García and Vlachos 2014
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†Titanochelon bacharidisi Vlachos et al. 2014
†Titanochelon bolivari Hernandez-Pacheco 1917
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Invalid names: Testudo richardi Bergounioux 1938 [synonym]
†Titanochelon ginsburgi Broin 1977
†Titanochelon kayadibiensis Karl et al. 2021
†Titanochelon perpiniana Depéret 1885
†Titanochelon punicum Arambourg 1979
†Titanochelon schafferi Szalai 1931
†Titanochelon schleichi Pappa et al. 2023
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| I. R. Swingland and M. W. Klemens 1989 | See Crumly (1984) | |
| E. Vlachos 2021 | The members of Geochelonini represented in the Greek fossil record can be easily distinguished from those belonging to Testudinini (see below) based on several characters from the shell: large size, with individuals that reach or exceed one meter in carapace length; absence of cervical scute; contact between marginal 6 and pleural 3; and absence of a plastral hinge. | |
| I. Pappa et al. 2023 | based on the alternating pattern of costal plates, the good lateral coincidence between the costo-peripheral sutures and pleuro-marginal sulci, the presence of a well-developed epiplastral lip, and the presence of four visible digits in the pes |