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Canis armbrusteri

Mammalia - Carnivora - Canidae

Taxonomy
Canis armbrusteri was named by Gidley (1913). Its type specimen is USNM 7662, a mandible, and it is a 3D body fossil. Its type locality is Cumberland Cave, which is in an Irvingtonian cave horizon in Maryland.

It was synonymized subjectively with Canis lupus by Martin (1974) and Miller (1976).

Sister species lacking formal opinion data

Synonymy list
YearName and author
1913Canis armbrusteri Gidley
1979Canis armbrusteri Nowak
1980Canis armbrusteri Kurten and Anderson p. 169
2008Canis armbrusteri Wang et al.
2009Canis armbrusteri Tedford et al. p. 137 figs. 40, 43, 44, 52, 53A–D, 54A–F, 55A–D; appendices 2–4
2019Canis armbrusteri Albright et al. p. 173

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
RankNameAuthor
Mammaliamorpha
Mammaliaformes
classMammalia
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Ferae()
CarnivoramorphaWyss and Flynn 1993
CarnivoraformesFlynn et al.
orderCarnivora
familyCanidae
subfamilyCaninaeGill 1872
genusCanis
speciesarmbrusteri

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Canis armbrusteri Gidley 1913
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Diagnosis
ReferenceDiagnosis
R. H. Tedford et al. 2009Canis armbrusteri shares with C. lupus and C. dirus the following synapomorphies: incisive foramina extend posterior to limit of canine alveoli, strong posterior expansion of paroccipital process, posterior expansion of frontal sinus to frontoparietal suture, and very reduced M1 parastyle lacking union with prepara- crista. It differs from C. lupus in that it retains premolars in which P3 and p2–p3 usually have posterior cusps, and the second posterior cusp of p4 is separate from the posterior cingulum. Like C. dirus, it also differs from C. lupus in having short-crowned and relatively straight canines, the metaconid of m1–m2 reduced, and a reduced P4 protocone. Although a sister taxon to C. dirus, C. armbrusteri lacks the other hyper- carnivorous dental features characteristic of C. dirus including a primitively retained labial cingulum on M1.