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Osmylidae (lance lacewing)

Insecta - Neuroptera - Osmylidae

Synonymy list
YearName and author
1815Osmylidae Leach
1955Osmylidae Riek p. 678
1992Osmylidae Carpenter
2008Osmylidae Engel and Grimaldi
2008Osmylidae Menon and Makarkin
2018Osmylidae Engel et al.
2019Osmylidae Winterton et al. p. 13

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
RankNameAuthor
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
superorderNeuropterida
—Metaxyneuroptera
—Eidoneuroptera
orderNeuroptera
—Euneuroptera
superfamilyOsmyloideaLeach 1815
familyOsmylidaeLeach 1815
familyOsmylidaeLeach 1815

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Osmylidae Leach 1815 [lance lacewing]
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G. †Archaeosmylidia Makarkin et al. 2014
+
†Archaeosmylidia fusca Makarkin et al. 2014
G. †Dimidiosmylus Khramov 2014
+
†Dimidiosmylus ramosus Khramov 2014
G. †Dzhidosmylus Khramov 2020
+
†Dzhidosmylus solus Khramov 2020
Subfm. Eidoporisminae Esben-Petersen 1917
+
G. †Stenosmylina Jepson et al. 2009
+
†Stenosmylina medialis Jepson et al. 2009
G. †Erlikosmylus Khramov 2014
+
†Erlikosmylus obscurus Khramov 2014
Subfm. Gumillinae Navas 1912
+
G. †Allotriosmylus Yang et al. 2010
+
†Allotriosmylus uniramosus Yang et al. 2010
G. †Enodinympha Ren and Engel 2007
+
†Enodinympha imperfecta Ren and Engel 2007
†Enodinympha translucida Ren and Engel 2007
G. †Epiosmylus Panfilov 1980
+
†Epiosmylus longicornis Panfilov 1980
†Epiosmylus longus Khramov 2017
†Epiosmylus panfilovi Ren and Yin 2002
G. Gumilla Navas 1912
G. †Kolbasinella Khramov 2014
+
†Kolbasinella elongata Khramov 2014
G. †Nilionympha Ren and Engel 2007
+
†Nilionympha pulchella Ren and Engel 2007
†Nilionympha shantouensis Li et al. 2018
G. †Nuddsia Menon and Makarkin 2008
+
†Nuddsia longiantennata Menon and Makarkin 2008
†Nuddsia magnifica Myskowiak et al. 2016
†Nuddsia repatriata Martins-Neto 2010
†Nuddsia simplex Li et al. 2024
Invalid names: Burmaleon Myskowiak et al. 2016 [synonym]
G. †Osmylochrysa Jepson et al. 2012
+
†Osmylochrysa anomala Jepson et al. 2012
†Osmylochrysa fragilis Jepson et al. 2012
†Osmylochrysa navasia Khramov 2020
G. †Tenuosmylus Wang et al. 2009
+
†Tenuosmylus brevineurus Wang et al. 2009
Invalid names: Epiosmylinae Panfilov 1980 [synonym]
G. †Karaosmylus Makarkin 1990
+
†Karaosmylus latus Panfilov 1980
Invalid names: Parosmylus Panfilov 1980 [replaced]
Subfm. Kempyninae Carpenter 1943
+
G. †Arbusella Khramov 2014
+
†Arbusella bella Khramov 2014
†Arbusella magna Khramov et al. 2017
†Arbusella platyptera Ma and Wang 2022
G. †Cretosmylus Makarkin 1990
+
†Cretosmylus sibiricus Makarkin 1990
G. †Euporismites Tillyard 1916
+
†Euporismites balli Tillyard 1916
G. †Jurakempynus Wang et al. 2011
+
†Jurakempynus arcanus Khramov 2014
†Jurakempynus bellatulus Wang et al. 2011
†Jurakempynus epunctatus Wang et al. 2011
†Jurakempynus loculosus Ma et al. 2020
†Jurakempynus sinensis Wang et al. 2011
†Jurakempynus sublimis Khramov 2014
G. †Kempynosmylus Makarkin 2014
+
†Kempynosmylus zherikhini Makarkin 2014
G. †Mirokempynus Ma et al. 2020
+
G. †Ponomarenkius Khramov et al. 2017
+
†Ponomarenkius excellens Khramov et al. 2017
G. †Sauktangida Khramov 2014
+
†Sauktangida aenigmatica Khramov 2014
G. †Kubekius Khramov 2017
+
†Kubekius multiramosus Khramov 2017
G. †Mesosmylidus Jepson et al. 2012
+
†Mesosmylidus vulgaris Jepson et al. 2012
Subfm. †Mesosmylininae Bode 1953
+
G. †Mesosmylina Bode 1953
+
†Mesosmylina angusta Khramov 2014
†Mesosmylina exornata Bode 1953
†Mesosmylina falcifera Ansorge 1996
†Mesosmylina mongolica Ponomarenko 1984
†Mesosmylina shurabica Khramov 2014
†Mesosmylina sibirica Ponomarenko 1985
Subfm. Osmylinae Leach 1815
+
G. †Lithosmylus Carpenter 1943
+
†Lithosmylus columbianus Cockerell 1908
G. Osmylus Latreille 1802
G. †Vetosmylus Ma et al. 2020
+
†Vetosmylus maculosus Ma et al. 2020
†Vetosmylus tentus Ma et al. 2020
G. †Osmylopsis Handlirsch 1906
+
†Osmylopsis duplicata Giebel 1856
+
Invalid names: Pterinoblattina binneyi Scudder 1886 [objective synonym]
G. †Palaeothyridosmylus Wang et al. 2009
+
G. †Petrushevskia Martynova 1958
+
†Petrushevskia borisi Martynova 1958
G. †Pronymphites Panfilov 1980
+
†Pronymphites elegans Panfilov 1980
Subfm. Protosmylinae Krüger 1913
+
G. †Juraheterosmylus Wang et al. 2010
+
†Juraheterosmylus antiquatus Wang et al. 2010
†Juraheterosmylus astictus Wang et al. 2010
†Juraheterosmylus minor Wang et al. 2010
G. †Jurosmylus Makarkin and Archibald 2005
+
†Jurosmylus atalantus Panfilov 1980
†Jurosmylus parvulus Khramov 2014
Invalid names: Mesosmylus Panfilov 1980 [replaced]
G. †Myanmarosmylus Li et al. 2024
+
†Myanmarosmylus wintertoni Li et al. 2024
G. †Osmylidia Cockerell 1908
+
†Osmylidia donnae Makarkin et al. 2021
†Osmylidia glastrai Makarkin et al. 2021
†Osmylidia picta Hagen 1856
†Osmylidia requieta Scudder 1890
†Osmylidia taliae Makarkin et al. 2021
Invalid names: Oligosmylus Krüger 1913 [objective synonym], Protosmylus Krüger 1913 [synonym]
G. †Protosmylina Jepson et al. 2009
+
†Protosmylina bifasciata Jepson et al. 2009
G. †Pseudosmylidia Makarkin 2017
+
†Pseudosmylidia relicta Makarkin 2017
G. †Sogjuta Martynova 1958
+
†Sogjuta shartegica Khramov 2011
†Sogjuta speciosa Martynova 1958
G. †Scapoptera Panfilov 1980
+
†Scapoptera recta Panfilov 1980
Subfm. Spilosmylinae Kruger 1913
+
G. †Ensiosmylus Khramov 2014
+
†Ensiosmylus acutus Khramov 2014
G. †Imanosmylus Makarkin 1994
+
†Imanosmylus ussuriensis Makarkin 1994
G. †Zakamnosmylus Khramov 2020
+
†Zakamnosmylus elongatus Khramov 2020
G. †Stenochrysa Jepson et al. 2012
+
†Stenochrysa gradata Jepson et al. 2012
G. †Tengriosmylus Khramov 2014
+
†Tengriosmylus magnificus Khramov 2014
G. †Tetanoptilon Bode 1953
+
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Sinemurian to the top of the Sarmatian or 199.30000 to 11.60800 Ma
Minimum age of oldest fossil (stem group age): 190.8 Ma

Collections (59 total)


Time interval Ma Country or state Original ID and collection number
Hettangian - Sinemurian201.3 - 190.8Kyrgyzstan (Tonskiy) Petrushevskia borisi, Sogjuta speciosa (105087)
Sinemurian199.3 - 190.8Kyrgyzstan (Tonskiy) Petrushevskia borisi (113412)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Mesosmylina exornata (105002) Tetanoptilon brunsvicense (107323)
Early/Lower Toarcian183.0 - 182.0Germany (Mecklenburg-Vorpommern) Mesosmylina falciferum (114993)
Toarcian182.7 - 174.1Kyrgyzstan (Batken) Erlikosmylus obscurus (118023) Mesosmylina shurabica (105003) Tengriosmylus magnificus, Mesosmylina angusta (110817)
Toarcian - Aalenian182.7 - 170.3Kyrgyzstan (Osh) Sauktangida aenigmatica (156499)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) Osmylidae indet., Mesosmylina sibirica, Kubekius multiramosus, Epiosmylus longus (118010)
Bathonian168.3 - 166.1Mongolia Mesosmylina mongolica (119621)
Callovian - Oxfordian166.1 - 157.3China (Nei Mongol) Allotriosmylus uniramosus, Archaeosmylidia fusca (113365) Enodinympha translucida, Nilionympha pulchella, Nilionympha imperfecta, Palaeothyridosmylus sp., Palaeothyridosmylus septemaculatus, Epiosmylus panfilovi, Tenuosmylus brevineurus (113367) Jurakempynus sinensis, Jurakempynus bellatulus, Jurakempynus epunctatus, Juraheterosmylus antiquatus, Juraheterosmylus astictus, Juraheterosmylus minor (113366) Mirokempynus profundobifurcus, Jurakempynus loculosus, Arbusella platyptera (219840) Nilionympha shantouensis (122589) Ponomarenkius sp., Ponomarenkius excellens, Arbusella magna (113371) Vetosmylus tentus, Vetosmylus maculosus (210302)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Arbusella sp. (106279) Ensiosmylus acutus (113439) Jurosmylus parvulus, Jurakempynus arcanus, Arbusella sp. (105970) Jurosmylus parvulus, Kolbasinella elongata, Arbusella bella (105426) Mesosmylus atalantus, Epiosmylus longicornis (106101) Pronymphites elegans, Parosmylus latus (105971) Scapoptera recta, Jurosmylus parvulus (106102)
Tithonian152.1 - 145.0Russian Federation (Chita) Mesosmylina sp. (125407)
Tithonian152.1 - 145.0Mongolia Jurakempynus sublimis (125978) Osmylidae indet., Sogjuta shartegica, Dimidiosmylus ramosus (104748)
Early/Lower Berriasian145.0 - 140.2United Kingdom (England) Orthoptera indet. (105966)
Late/Upper Berriasian145.0 - 140.2United Kingdom (England) Mesosmylidus vulgaris, Osmylochrysa anomala, Osmylochrysa fragilis, Stenochrysa gradata (122909)
Late/Upper Hauterivian136.4 - 130.0United Kingdom (England) Stenosmylina medialis (106417)
Early/Lower Barremian130.0 - 125.45United Kingdom (England) Protosmylina bifasciata (118921)
Aptian125.0 - 113.0Russian Federation (Buryatia) Cretosmylus sibiricus (119498) Kempynosmylus zherikhini (106103) Osmylochrysa navasia, Zakamnosmylus elongatus, Dzhidosmylus solus (113350)
Late/Upper Aptian122.46 - 113.0Brazil (Ceara) Nuddsia repatriata (34901)
Late/Upper Aptian122.46 - 113.0Brazil (Ceará) Nuddsia longiantennata (122557)
Early/Lower Cenomanian99.6 - 93.5Myanmar Burmaleon magnificus, Nuddsia simplex (161726) Nuddsia simplex (212006) Nuddsia simplex, Myanmarosmylus wintertoni (196491) Osmylidae indet. (111688)
Maastrichtian72.1 - 66.0Russian Federation (Primorskiy) Imanosmylus ussuriensis (119699)
Ypresian56.0 - 47.8Australia (Queensland) Euporismites balli (111780 125666 139941)
Ypresian56.0 - 47.8Canada (British Columbia) Osmylidia donnae (31677) Osmylidia sp. (104793)
Ypresian56.0 - 47.8USA (Washington) Osmylidia glastrai (103206)
Bridgerian50.3 - 46.2USA (Colorado) Osmylidia taliae (181561)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Osmylidia picta (110735)
Priabonian38.0 - 33.9Poland Osmylus pictus (120811)
Chadronian37.2 - 33.9USA (Colorado) Lithosmylus columbianus (151548) Osmylidia requieta (113749 124624) Osmylidia requietus (106379) Osmylus columbianus (106453) Osmylus requietus (106384) Pseudosmylidia relicta (106381)
Sarmatian12.7 - 11.608Romania Osmylus sp. (156756)