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Harknessellidae

Rhynchonellata - Orthida - Harknessellidae

Taxonomy
Harknessellidae was named by Bancroft (1928). Its type is Harknessella.

It was assigned to Dalmanellacea by Wright (1964); to Enteletacea by Hints (1975); and to Dalmanelloidea by Williams et al. (2000).

Synonymy list
YearName and author
1928Harknessellidae Bancroft
1964Harknessellidae Wright
1975Harknessellidae Hints
2000Harknessellidae Williams et al. p. 800

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Spiralia
superphylumLophotrochozoa
—Lophophorata
—PanbrachiopodaCarlson and Cohen 2020
RankNameAuthor
phylumBrachiopodaCuvier 1805
subphylumRhynchonelliformeaWilliams et al. 1996
classRhynchonellataWilliams et al. 1996
orderOrthidaSchuchert and Cooper 1932
suborderDalmanellidinaMoore 1952
superfamilyDalmanelloideaSchuchert 1913
familyHarknessellidaeBancroft 1928
familyHarknessellidaeBancroft 1928

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Harknessellidae Bancroft 1928
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G. †Haymina Bogoyavlenskaya 1991
G. †Horderleyella Bancroft 1928
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†Horderleyella chacaltanai Villas et al. 2015
†Horderleyella fragilis Bergström 1968
G. †Isabelella Reyes-Abril et al. 2013
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†Isabelella fascicostellata Reyes-Abril et al. 2013
G. †Kampella Baarli 1988
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†Kampella guttula Baarli 1988
G. †Reuschella Bancroft 1928
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†Reuschella americana Cooper 1956
†Reuschella herreraensis Villas 1992
†Reuschella horderleyensis Bancroft 1929
†Reuschella inexpectata Temple 1968
†Reuschella magna Hints 1975
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitep
Environment: marineuc
Locomotion: stationaryo
Attached: yeso
Life habit: epifaunalo
Diet: suspension feedero
Vision: blindc
Created: 2004-02-28 14:11:06
Modified: 2009-04-11 14:18:14
Source: o = order, c = class, p = phylum, uc = unranked clade
References: Aberhan et al. 2004, Nesnidal et al. 2013

Age range: base of the Dapingian to the top of the Telychian or 471.30000 to 432.90000 Ma

Collections (201 total)


Time interval Ma Country or state Original ID and collection number
Dapingian471.3 - 469.4China Horderleyella sp. (232225)
Darriwilian469.4 - 458.2China (Sichuan) Horderleyella rude (64229 64235 64778 64782 203293)
Early/Lower Llanvirn468.0 - 461.1China (Yunnan) Harknessella sp. (5427)
Llanvirn468.0 - 458.2United Kingdom (Shropshire) Horderleyella sp. (45052)
Llanvirn468.0 - 458.2Russian Federation Haymina carinata (219874)
Early/Lower Llandeilo461.1 - 458.2United Kingdom (Wales) Horderleyella sp. (3366)
Late/Upper Llanvirn461.1 - 458.2United Kingdom (Wales) Horderleyella sp. (69532 69571 69581 69582 69583)
Late/Upper Llanvirn461.1 - 458.2United Kingdom Horderleyella convexa (69563 69564 69565 69597 69598 69601) Horderleyella sp. (69533 69534 69566 69567 69568 69569 69570 69572 69573 69574 69575 69576 69577 69578 69580)
Late/Upper Llanvirn461.1 - 458.2China (JiangSu) Horderleyella sp. (7938)
Llandeilo461.1 - 458.2United Kingdom Horderleyella convexa (69593 69596)
Dobrotivian461.0 - 459.7Spain (Cuidad Real) Isabelella fascicostellata (219875 219876)
Dobrotivian461.0 - 459.7Spain (Toledo and Cuidad Real) Horderleyella sp. (20893)
Berounian - Vinice459.7 - 457.3Spain (Zaragoza) Horderleyella sp., Reuschella herreraensis (21089) Reuschella herreraensis (166737)
Berounian459.7 - 450.1Spain (Cuidad Real) Reuschella herreraensis (21099)
Berounian - Caradoc459.7 - 449.5Portugal Horderleyella sp. (20954)
Bohdalec459.3 - 445.2Czechia Horderleyella bouceki (3872)
Zahorany - Late/Upper Ordovician459.3 - 443.1Czechia Horderleyella bouceki, Reuschella macrostoma (3865)
Costonian458.2 - 457.2United Kingdom (Wales) Horderleyella sp. (188595)
Costonian458.2 - 457.2United Kingdom (Shropshire) Harknessella subplicata, Horderleyella plicata (45279) Horderleyella plicata (45278)
Costonian - Harnagian458.2 - 457.2United Kingdom (Wales) Reuschella horderleyensis (188594)
Harnagian458.2 - 457.2United Kingdom (England) Smeathenella harnagensi (174723)
Harnagian458.2 - 457.2United Kingdom (Shropshire) Horderleyella sp. (45323)
Harnagian458.2 - 457.2United Kingdom (Wales) Horderleyella sp. (43634) Reuschella horderleyensis (43635)
Soudleyan458.2 - 457.2United Kingdom (Wales) Reuschella horderleyensis (3301 3362 43636 43637 43638 43639 188588 188589 188593 188596 192387)
Soudleyan458.2 - 457.2United Kingdom (England) Reuschella oblonga, Reuschella horderleyensis (3302 3303)
Soudleyan458.2 - 457.2United Kingdom (Shropshire) Reuschella horderleyensis (46883 46885 46886 46887 46891 46892 46893)
Aurelucian458.2 - 456.6United Kingdom (Shropshire) Smeathenella strophomenoides (219878)
Soudleyan - Marshbrookian458.2 - 454.0United Kingdom (England) Horderleyella sp., Reuschella sp. (3654) Reuschella sp. (3649 3651)
Soudleyan - Marshbrookian458.2 - 454.0United Kingdom (Wales) Reuschella sp. (3368 3369 3370 3372 3373)
Sandbian458.2 - 452.8Estonia Horderleyella kegelensis (164219 164222 164248) Horderleyella sp. (164258) Horderleyella sp., Horderleyella kegelensis (164246 164247 164257)
Caradoc458.2 - 449.5United Kingdom (England ) Harknessella sp. (434)
Turinian457.3 - 453.8USA (Virginia) Reuschella americana (164197) Reuschella edsoni (76338) Reuschella sp. (76449)
Longvillian457.2 - 455.0United Kingdom (Wales) Reuschella horderleyensis (43640 192388 192389)
Longvillian - Marshbrookian457.2 - 454.0Norway Horderleyella kegelensis (2763)
Woolstonian457.2 - 454.0United Kingdom (Wales) Reuschella sp. (3305)
Burrellian456.6 - 452.5United Kingdom (Scotland) Reuschella americana (165050)
Burrellian456.6 - 452.5Peru Horderleyella chacaltanai (167133 167134)
Actonian - Onnian454.0 - 449.5United Kingdom (England) Reuschella sp. (3650 3655 3656 3658)
Keila453.9 - 452.8Estonia (Lääne-Viru) Dalmanella kegelensis (174056) Horderleyella kegelensis (197525) Horderleyella oanduensis (161216)
Keila453.9 - 452.8Latvia Horderleyella kegelensis (207211)
Keila453.9 - 452.8Russian Federation (Novgorod oblast) Horderleyella kegelensis (235850)
Keila453.9 - 452.8Lithuania Horderleyella kegelensis (195479 207210 207239) Horderleyella sp. (161228 192436)
Keila453.9 - 452.8Russian Federation Dalmanella kegelensis (173853 173854 173902 173903 173911 173914 173930 173961 173982 174013 174015 174052) Horderleyella kegelensis (235843)
Keila453.9 - 452.8Estonia (Harju) Horderleyella kegelensis (195554 197527 197528)
Keila453.9 - 452.8Estonia Dalmanella kegelensis (166045 173859) Horderleyella kegelensis (161153 161237 161280 197517 197518 197519 197521) Horderleyella oanduensis (161138) Horderleyella sp. (168964 168981 197549)
Keila453.9 - 452.8Estonia (Lääne) Horderleyella kegelensis (195653 195654)
Keila453.9 - 452.8Estonia (Hiiu) Horderleyella kegelensis (161182)
Onnian - Cautleyan453.0 - 448.8Belgium Reuschella sp. (185313)
Oandu452.8 - 451.8Russian Federation Reuschella magna (161371)
Oandu452.8 - 451.8Estonia Reuschella magna (161116 161218)
Oandu452.8 - 451.8Russian Federation (Leningrad oblast) Horderleyella kegelensis (235821)
Oandu452.8 - 451.8Lithuania Horderleyella kegelensis (192878 192879 195476) Reuschella magna (161167 161229 192438 207241)
Johvi452.8 - 449.9Latvia Horderleyella kegelensis (207209)
Johvi452.8 - 449.9Lithuania Horderleyella kegelensis (207208)
Katian452.8 - 445.2Estonia Horderleyella sp. (164264 164266)
Cheneyan452.5 - 451.3United Kingdom (Scotland) Reuschella americana (164412)
Cautleyan449.2 - 448.8United Kingdom (Wales) Harknessella stevensorum (185102)
Cautleyan449.2 - 448.8Ireland (Leinster) Reuschella sp. (89130)
Cautleyan - Rawtheyan449.2 - 445.2United Kingdom (Wales) Reuschella sp. (8290 8291)
Rawtheyan448.8 - 445.2United Kingdom (England) Reuschella sp. (3667)
Rawtheyan448.8 - 445.2United Kingdom (Wales) Harknessella stevensorum (185103)
Hirnantian445.2 - 443.1Sweden Horderleyella sp. (2768)
Hirnantian445.2 - 443.1United Kingdom (England) Reuschella inexpectata (80657)
Hirnantian445.2 - 443.1Argentina Reuschella sp. (8890 8891)
Hirnantian445.2 - 443.1China (ZheJiang) Reuschella sp. (7775)
Hirnantian445.2 - 443.1United Kingdom (Wales) Reuschella sp. (8405)
Hirnantian445.2 - 443.1Sweden (Dalarne) Horderleyella fragilis (73455)
Hirnantian445.2 - 443.1China (Hubei) Horderleyella sp. (25816)
Kosovian445.2 - 443.1Morocco Horderleyella sp. (21002)
Kosovian445.2 - 443.1Portugal Horderleyella sp. (20934)
Porkuni445.2 - 443.1Sweden (Dalarne) Horderleyella fragilis (152984 153018)
Porkuni445.2 - 443.1Estonia (Lääne-Virumaa) Reuschella sp. (80715 152014 152015 228451)
Rhuddanian443.1 - 440.5Norway (Asker) Reuschella sp. (160131 191023 191025 191026 191027 191028 191029 191030)
Rhuddanian443.1 - 440.5United Kingdom (Wales) Reuschella inexpectata (8407)
Llandovery443.1 - 432.9Norway Reuschella sp. (502)
Telychian438.6 - 432.9Norway (Sandvika) Kampella guttula (160115 160116)