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Panorthoptera

Insecta

Synonymy list
YearName and author
1928Panorthoptera Crampton
2004Panorthoptera Béthoux et al.
2019Panorthoptera Schubnel et al. p. 500

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
RankNameAuthor
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—PolyneopteraMartynov 1938
superorderArchaeorthoptera
—PanorthopteraCrampton 1928
—PanorthopteraCrampton 1928

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Panorthoptera Crampton 1928
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G. †Bruaylogus Coty et al. 2014
+
†Bruaylogus magnificus Coty et al. 2014
Fm. †Cacurgidae Handlirsch 1911
+
G. †Cacurgus Handlirsch 1911
+
†Cacurgus avionensis Schubnel et al. 2019
†Cacurgus spilopterus Handlirsch 1911
G. †Heterologopsis Brauckmann and Koch 1982
+
†Heterologopsis ruhrensis Brauckmann and Koch 1982
G. †Spilomastax Handlirsch 1911
+
†Spilomastax oligoneurus Handlirsch 1911
Or. †Caloneurodea Martynov 1938
+
Fm. †Apsidoneuridae Carpenter 1961
+
G. †Apsidoneura Carpenter 1943
+
†Apsidoneura flexa Carpenter 1943
G. †Homaloptila Handlirsch 1919
+
†Homaloptila similis Meunier 1911
G. †Sinaspidoneura Huang et al. 2020
+
†Sinaspidoneura magnifica Huang et al. 2020
G. †Aviogramma Prokop et al. 2014
+
†Aviogramma gracilis Prokop et al. 2014
Fm. †Caloneuridae Handlirsch 1906
+
G. †Caloneura Brongniart 1885
+
†Caloneura dawsoni Brongniart 1885
+
Invalid names: Caloneura longicornis Handlirsch 1919 [synonym], Caloneura major Handlirsch 1919 [synonym], Caloneura picta Handlirsch 1919 [synonym], Caloneura robusta Handlirsch 1919 [synonym], Caloneura subtilis Bolton 1925 [synonym], Homaloneura royeri Meunier 1911 [synonym]
†Caloneura dawsonii Brongniart 1885
Invalid names: Boltonaloneura Rasnitsyn 2002 [synonym], Confusio Handlirsch 1919 [synonym]
G. †Caloneurella Carpenter 1934
+
†Caloneurella carbonaria Carpenter 1934
G. †Eohymen Martynov 1937
+
†Eohymen maculipennis Martynov 1937
G. †Euthygramma Martynov 1928
+
†Euthygramma parallellum Martynov 1928
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Invalid names: Euthygramma aberrans Martynov 1938 [synonym], Euthygramma curvatum Martynov 1931 [synonym]
†Euthygramma storozhenkoi Rasnitsyn 2025
Invalid names: Paleuthygrammella Engel 1995 [synonym], Pseudogramma Carpenter 1943 [synonym]
G. †Isadistica Rasnitsyn and Aristov 2013
+
†Isadistica issada Rasnitsyn and Aristov 2013
†Isadistica longa Rasnitsyn and Aristov 2013
G. †Ligogramma Béthoux et al. 2004
+
†Ligogramma sinuosa Béthoux et al. 2004
†Ligogramma wichita Beckemeyer 2009
G. †Pleisiogramma Carpenter 1943
+
†Pleisiogramma maueri Novokshonov 1998
†Pleisiogramma medialis Carpenter 1943
+
Invalid names: Pleisiogramma reducta Carpenter 1943 [synonym]
G. †Vilviopsis Martynov 1938
+
†Vilviopsis extensa Martynov 1938
†Vilviopsis picta Rasnitsyn 2004
Invalid names: Amboneuridae Carpenter 1980 [synonym], Anomalogrammatidae Carpenter 1943 [synonym], Eohymenidae Martynov 1937 [synonym], Euthygrammatidae Martynov 1928 [synonym], Paleuthygrammatidae Carpenter 1943 [synonym], Plesiogrammatidae Carpenter 1943 [synonym], Sthenaroceridae Handlirsch 1906 [synonym]
G. †Fusiogramma Boderau et al. 2024
+
†Fusiogramma minuta Boderau et al. 2024
G. †Gallogramma Garrouste et al. 2017
+
†Gallogramma galadrieli Garrouste et al. 2017
G. †Gigagramma Béthoux et al. 2004
+
†Gigagramma carpenteri Béthoux et al. 2004
Fm. †Hapalopteridae Handlirsch 1906
+
G. †Aenigmatodes Handlirsch 1906
+
†Aenigmatodes danielsi Handlirsch 1906
G. †Arroyopteron Rasnitsyn 2004
+
†Arroyopteron indolatum Rasnitsyn 2004
G. †Carrizarroyo Rasnitsyn 2004
+
†Carrizarroyo calopterus Rasnitsyn 2004
G. †Carrizopteron Rasnitsyn 2004
+
†Carrizopteron longum Rasnitsyn 2004
G. †Endoiasmus Handlirsch 1906
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†Endoiasmus reticulatus Handlirsch 1906
G. †Geroneura Matthew 1889
+
†Geroneura ovata Bolton 1921
†Geroneura wilsoni Matthew 1889
G. †Hapaloptera Handlirsch 1906
+
†Hapaloptera gracilis Handlirsch 1906
†Hapaloptera obovata Bolton 1934
†Hapaloptera parallela Bolton 1934
G. †Protokollaria Brongniart 1885
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†Protokollaria ingens Brongniart 1885
G. †Spiculum Brauckmann and Herd 2005
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†Spiculum mendicum Brauckmann and Herd 2005
G. †Tshecalculus Novokshonov 2000
+
†Tshecalculus inaspectus Novokshonov 2000
Invalid names: Aenigmatodidae Handlirsch 1906 [synonym], Protokollariidae Handlirsch 1906 [synonym]
G. †Jarmiloptera Prokop et al. 2015
+
†Jarmiloptera mouralensis Prokop et al. 2015
Fm. †Lodevogrammatidae Nel et al. 2023
+
G. †Lodevogramma Nel et al. 2023
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†Lodevogramma pumilia Nel et al. 2023
G. †Lusitaneura Loureiro et al. 2010
+
†Lusitaneura covensis Loureiro et al. 2010
G. †Nanogramma Béthoux et al. 2004
+
†Nanogramma gandi Béthoux et al. 2004
G. †Oudardgramma Nel and Roques 2021
+
†Oudardgramma bruayensis Nel and Roques 2021
G. †Paleuthygramma Martynov 1930
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†Paleuthygramma acuta Carpenter 1943
+
Invalid names: Apsidoneura reducta Zimmerman 1962 [synonym]
†Paleuthygramma sharovi Béthoux et al. 2004
†Paleuthygramma sottyi Burnham 1984
†Paleuthygramma tenuicorne Martynov 1930
†Paleuthygramma ventricosa Rasnitsyn 2004
G. †Paraspiculum Dvořák et al. 2021
+
†Paraspiculum brauckmanni Dvořák et al. 2021
Fm. †Permobiellidae Tillyard 1937
+
G. †Aviobiella Nel and Roques 2021
+
†Aviobiella garroustei Nel and Roques 2021
G. †Permobiella Tillyard 1937
+
†Permobiella perspicua Tillyard 1937
G. †Pseudobiella Carpenter 1970
+
†Pseudobiella fasciata Carpenter 1970
G. †Pruvostiella Handlirsch 1922
+
†Pruvostiella lecomtei Pruvost 1919
Invalid names: Euthyneura Pruvost 1919 [replaced]
G. †Wappleria Santos et al. 2024
+
†Wappleria tremoris Santos et al. 2024
Invalid names: Hapalopteroidea Handlirsch 1906 [synonym], Pleisiogramma kemerovoense Martynova 1952 [nomen dubium], Stenaroceridae [empty]
Fm. †Geraridae Scudder 1885
+
G. †Anepitedius Handlirsch 1911
+
†Anepitedius giraffa Handlirsch 1911
G. †Axiologus Handlirsch 1906
+
†Axiologus thoracicus Handlirsch 1906
G. †Cantabrala Kukalová-Peck and Brauckmann 1992
+
†Cantabrala gandli Kukalová-Peck and Brauckmann 1992
G. †Genentomum Scudder 1885
+
†Genentomum subacutum Bolton 1911
†Genentomum validum Scudder 1885
+
Invalid names: Genentomum carri Cockerell 1917 [synonym]
G. †Gerarulus Handlirsch 1911
+
†Gerarulus radialis Handlirsch 1911
G. †Gerarus Scudder 1885
+
†Gerarus collaris Handlirsch 1911
+
Invalid names: Gerarus longicollis Handlirsch 1911 [synonym]
†Gerarus constrictus Scudder 1885
+
Invalid names: Gerarus angustus Handlirsch 1906 [synonym], Gerarus latus Handlirsch 1911 [synonym], Gerarus longus Handlirsch 1906 [synonym], Gerarus reductus Handlirsch 1911 [synonym], Rossites inopinus Richardson 1956 [synonym]
†Gerarus danielsi Handlirsch 1906
†Gerarus fischeri Brongniart 1885
+
Invalid names: Archaeacridites bruesi Meunier 1909 [synonym], Archaeacridites elegantissima Meunier 1909 [synonym], Sthenaropoda agnusi Lameere 1917 [synonym], Sthenaropoda lerichei Lameere 1917 [synonym]
†Gerarus mazonus Scudder 1885
†Gerarus symmetricus Bolton 1934
†Gerarus teutonicus Brauckmann and Herd 2005
†Gerarus vetus Scudder 1885
Invalid names: Archaeacridites Meunier 1909 [synonym], Genopteryx Scudder 1885 [synonym], Rossites Richardson 1956 [synonym], Sthenaropoda Brongniart 1885 [synonym]
G. †Hirsutgerarus Zessin 2012
+
†Hirsutgerarus saxonicus Zessin 2012
G. †Nacekomia Richardson 1956
+
†Nacekomia rossae Richardson 1956
G. †Osnogerarus Kukalová-Peck and Brauckmann 1992
+
†Osnogerarus trecwithiensis Kukalová-Peck and Brauckmann 1992
G. †Owadpteron Dvořák et al. 2019
+
†Owadpteron dareki Dvořák et al. 2019
G. †Parapalaeomastax Dvořák et al. 2019
+
†Parapalaeomastax dariuszi Dvořák et al. 2019
G. †Ploetzgerarus Zessin 2009
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†Ploetzgerarus krempieni Zessin 2009
G. †Progenentomum Handlirsch 1906
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†Progenentomum carbonis Handlirsch 1906
Invalid names: Genopterygidae Richardson 1956 [synonym], Sthenaropodidae Handlirsch 1906 [synonym]
Fm. †Permostridulidae Béthoux et al. 2003
+
G. †Permostridulus Béthoux et al. 2003
+
†Permostridulus brongniarti Béthoux et al. 2003
G. †Sinogerarus Gu et al. 2017
+
†Sinogerarus pectinatus Gu et al. 2017
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range: base of the Marsdenian to the top of the Changhsingian or 321.30000 to 251.90200 Ma

Collections (84 total)


Time interval Ma Country or state Original ID and collection number
Marsdenian321.3 - 320.0Germany (Ruhr) Heterologopsis ruhrensis (155261)
Westphalian A318.7 - 316.9Canada (Nova Scotia) Caloneurodea indet. (216802)
Westphalian A318.7 - 316.9United Kingdom (England) Genentomum subacutum (127291)
Westphalian A318.7 - 316.9Poland Owadpteron dareki, Parapalaeomastax dariuszi (107805)
Westphalian A318.7 - 316.9Canada (New Brunswick) Geroneura wilsoni (114468)
Westphalian A318.7 - 316.9France Oudardgramma bruayensis (122506)
Westphalian B316.9 - 314.6United Kingdom (Staffordshire) Geroneura ovata (34906)
Westphalian B316.9 - 314.6United Kingdom (Wales) Gerarus symmetricus, Hapaloptera obovata (151398) Hapaloptera parallela (151404)
Westphalian B - Westphalian C316.9 - 309.8China (Ningxia) Sinogerarus pectinatus (139576)
Westphalian C - Westphalian D314.6 - 307.4USA (Pennsylvania) Hapaloptera gracilis (114456)
Westphalian C - Westphalian D314.6 - 307.4France (Nord Pas de Calais) Bruaylogus magnificus (153462)
Westphalian C - Westphalian D314.6 - 307.4France Aviogramma gracilis, Cacurgus avionensis, Aviobiella garroustei, Fusiogramma minuta (122504)
Westphalian D309.8 - 307.4USA (Illinois) Aenigmatodes danielsi, Gerarus danielsi, Progenentomum carbonis (146857) Genentomum carri (122527) Gerarus collaris, Genopteryx constricta, Genentomum validum, Gerarus vetus, Gerarus mazonus, Gerarus longus, Gerarus angustus, Axiologus thoracicus, Endoiasmus reticulatus (21981) Gerarus danielsi (122512) Gerarus latus, Gerarus collaris, Gerarus reductus, Gerarus longicollis, Gerarulus radialis, Anepitedius giraffa, Cacurgus spilopterus, Spilomastax oligoneurus (112630) Gerarus vetus (151855) Gerarus vetus, Gerarus danielsi (151848 151849) Gerarus vetus, Gerarus danielsi, Nacekomia rossae (112638) Rossites inopinus (122530 122531)
Westphalian D309.8 - 307.4United Kingdom (Somerset) Gerarus danielsi (156594)
Westphalian D309.8 - 307.4France (Nord-Pas-de-Calais) Euthyneura lecomtei (124902)
Westphalian D309.8 - 307.4Germany (Lower Saxony) Caloneurodea indet., Geraridae indet., Spiculum mendicum (122515) Gerarus teutonicus, Primakollaria amicorum (122518) Osnogerarus trecwithiensis (126589) Paraspiculum brauckmanni (224159)
Westphalian D309.8 - 307.4USA (Pennsylvania) Amboneura klosei (122502) Caloneurella carbonaria (124504) Caloneurodea indet. (114649)
Cantabrian307.4 - 305.0Spain Cantabrala gandli (126590)
Kasimovian307.0 - 303.7USA (New Mexico) Pseudobiella fasciata (49663)
Gzhelian303.7 - 298.9Spain Wappleria tremoris (239851)
Gzhelian303.7 - 298.9France Caloneura dawsoni, Caloneura subtilis (124916) Gerarus fischeri, Sthenaropoda lerichei, Archaeacridites bruesi, Sthenarocera pachytyloides, Homaloneura similis, Homaloneura royeri, Archaeacridites elegantissima, Sthenaropoda fischeri (121979) Sthenaropoda fischeri, Sthenaropoda lerichei, Archaeacridites bruesi, Protokollaria ingens, Sthenarocera pachytyloides, Sthenarocera bureaui, Caloneura picta, Caloneura major, Caloneura longicornis, Caloneura robusta, Caloneura dawsonii (103635)
Stephanian B301.4 - 300.9France Apsidoneura sottyi (122490)
Stephanian C300.9 - 300.3Germany Hirsutgerarus saxonicus (163476) Ploetzgerarus krempieni (122505)
Stephanian C300.9 - 300.3Portugal Lusitaneura covensis (103240)
Stephanian C300.9 - 300.3Germany (Saxony-Anhalt) Laspeyresia wettinensis (122578)
Asselian298.9 - 293.52Germany Panorthoptera indet. (222678)
Asselian298.9 - 293.52USA (New Mexico) Paleuthygramma ventricosa, Carrizarroyo calopterus (124386) Vilviopsis picta (124385) Vilviopsis picta, Carrizopteron longum, Arroyopteron indolatum (124387)
Artinskian290.1 - 283.3Russian Federation (Perm) Vilvia densinervosa (153669)
Artinskian290.1 - 283.3France (Languedoc) Nanogramma gandi, Jarmiloptera mouralensis (105044) Paleuthygramma acuta (192422) Panorthoptera indet. (174411)
Artinskian290.1 - 283.3USA (Oklahoma) Apsidoneura reducta (104826) Ligogramma wichita (104821) Paleuthygramma acuta, Apsidoneura flexa, Anomalogramma parva, Gigagramma carpenteri, Ligogramma sinuosa (107405)
Artinskian290.1 - 283.3France Jarmiloptera mouralensis (174409)
Artinskian290.1 - 283.3USA (Kansas) Anomalogramma parva (114151) Caloneurodea indet. (114450) Caloneurodea indet., Permobiella perspicua, Paleuthygramma acuta, Apsidoneura flexa (111320) Paleuthygramma acuta (114152) Paleuthygramma acuta, Apsidoneura flexa, Plesiogramma medialis (111319) Permobiella perspicua, Paleuthygramma acuta (111219) Plesiogramma reducta (111217)
Artinskian - Wordian290.1 - 264.28France (Languedoc) Gallogramma galadrieli (192423)
Kungurian283.3 - 274.4Russian Federation (Suksun) Paleuthygramma tenuicornis (157042) Tshecalculus inaspectus, Paleuthygramma tenuicorne (108206)
Kungurian283.3 - 274.4Russian Federation Pleisiogramma maueri (107787)
Kungurian283.3 - 274.4France (Languedoc) Permostridulus brongniarti (104990 105053 122920)
Kungurian283.3 - 274.4France Lodevogramma pumilia, Caloneurodea indet. (103637)
Roadian274.4 - 266.9Russian Federation (Kazan) Euthygramma curvatum (116803)
Roadian274.4 - 266.9Russian Federation Euthygramma parallellum (116935)
Roadian274.4 - 266.9Russian Federation (Arkangelsk) Euthygramma aberrans (104993) Euthygramma parallellum (104716 116535 116933) Euthygramma parallellum, Vilviopsis extensa (116934) Paleuthygramma sharovi, Euthygramma parallelum (104992)
Roadian274.4 - 266.9Russian Federation (Kemerovo) Pleisiogramma kemerovoense (124605)
Capitanian264.28 - 259.51Russian Federation (Orenburg) Eohymen maculipennis (116607)
Capitanian264.28 - 259.51China (Anhui) Sinaspidoneura magnifica (103239)
Wuchiapingian259.51 - 254.14Russian Federation (Vologda) Isadistica issada, Isadistica longa (104754)
Changhsingian254.14 - 251.902Russian Federation (Orenburg) Euthygramma storozhenkoi (233223)