PBDB Taxon

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Protomyrmeleon

Insecta - Protomyrmeleontidae

Taxonomy

Protomyrmeleon was named by Geinitz (1888). It is the type genus of Protomyrmeleonidae, Protomyrmeleontoidea, Protomyrmeleontinae.

It was assigned to Protomyrmeleonidae by Handlirsch (1906), Bode (1953) and Henrotay et al. (1997); to Protomyrmeleontidae by Tillyard (1925), Martynov (1927), Zessin (1991), Carpenter (1992), Nel and Henrotay (1992), Ansorge (1996), Nel and Jarzembowski (1998), Nel et al. (2005) and Pritykina and Vassilenko (2014); and to Protomyrmeleontinae by Deregnaucourt et al. (2019).

Species

P. anglicanus, P. angustovenosus, P. brunonis (syn. P. bascharagensis), P. cretacicus, P. grandis, P. grasselensis, P. handlirschi, P. jurassicus, P. karatauensis, P. kazakhstanensis, P. pascholdi, P. pumilio, P. quadriordinum

Synonyms

Protomyrmeleon (Tillyardagrion) was named by Martynov (1927).

It was reranked as Tillyardagrion by Zessin (1991) and Carpenter (1992); it was synonymized subjectively with Protomyrmeleon by Nel and Henrotay (1992) and Henrotay et al. (1997).

It was assigned to Protomyrmeleon by Martynov (1927) and Bode (1953); and to Protomyrmeleontidae by Zessin (1991) and Carpenter (1992).
Zirzipanagrion was named by Zessin (1991).

It was synonymized subjectively with Protomyrmeleon by Nel et al. (2005).

It was assigned to Protomyrmeleontidae by Zessin (1991) and Nel and Henrotay (1992); and to Protomyrmeleonidae by Henrotay et al. (1997).

Synonymy list

Year	Name and author
1888	Protomyrmeleon Geinitz p. 204
1906	Protomyrmeleon Handlirsch p. 471
1925	Protomyrmeleon Tillyard p. 36
1927	Protomyrmeleon Martynov p. 759
1927	Protomyrmeleon (Tillyardagrion) Martynov p. 762
1953	Protomyrmeleon Bode
1953	Protomyrmeleon (Tillyardagrion) Bode
1991	Zirzipanagrion Zessin
1991	Protomyrmeleon Zessin p. 102
1991	Tillyardagrion Zessin p. 102
1991	Zirzipanagrion Zessin p. 112
1992	Protomyrmeleon Carpenter p. 68
1992	Tillyardagrion Carpenter p. 68
1992	Protomyrmeleon Nel and Henrotay p. 3
1992	Zirzipanagrion Nel and Henrotay p. 19
1996	Protomyrmeleon Ansorge p. 13
1997	Zirzipanagrion Henrotay et al. p. 397
1997	Protomyrmeleon Henrotay et al. p. 398
1998	Protomyrmeleon Nel and Jarzembowski p. 393
2005	Protomyrmeleon Nel et al. p. 192
2014	Protomyrmeleon Pritykina and Vassilenko p. 1642
2019	Protomyrmeleon Deregnaucourt et al. p. 310

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
infraclass	Hydropalaeoptera	Rohdendorf 1969
—	Euhydropalaeoptera
—	Odonatoptera	Martynov 1932
—	Palaeodonatoptera

Rank	Name	Author
—	Plesiodonatoptera
—	Apodonatoptera
—	Paneodonatoptera
—	Neodonatoptera
—	Euodonatoptera
—	Odonatoclada	Bechly 1999
—	Nodialata
—	Discoidalia
—	Stigmoptera
suborder	Archizygoptera
superfamily	Protomyrmeleontoidea	Handlirsch 1906
—	Terskejoptera
—	Moltenoptera
family	Protomyrmeleontidae	()
subfamily	Protomyrmeleontinae
genus	Protomyrmeleon

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Protomyrmeleon Geinitz 1888

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†Protomyrmeleon anglicanus Tillyard 1925

†Protomyrmeleon angustovenosus Martynov 1927

†Protomyrmeleon brunonis Geinitz 1888

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Invalid names: Protomyrmeleon bascharagensis Nel and Henrotay 1992 [synonym]

†Protomyrmeleon cretacicus Nel and Jarzembowski 1998

†Protomyrmeleon grandis Nel et al. 2005

†Protomyrmeleon grasselensis Bode 1953

†Protomyrmeleon handlirschi Martynov 1927

†Protomyrmeleon jurassicus Nel 1991

†Protomyrmeleon karatauensis Nel et al. 2005

†Protomyrmeleon kazakhstanensis Nel et al. 2005

†Protomyrmeleon pascholdi Zessin 1991

†Protomyrmeleon pumilio Nel et al. 2005

†Protomyrmeleon quadriordinum Zessin 1991

Invalid names: Tillyardagrion Martynov 1927 [synonym], Zirzipanagrion Zessin 1991 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range: base of the Rhaetian to the top of the Late/Upper Hauterivian or 208.50000 to 130.00000 Ma

Collections (17 total)

Time interval	Ma	Country or state	Original ID and collection number
Rhaetian	208.5 - 201.3	United Kingdom (England)	P. anglicanus (115222)
Sinemurian	199.3 - 190.8	United Kingdom (England)	P. brunonis (192973)
Early/Lower Toarcian	183.0 - 182.0	Germany (Mecklenburg-Vorpommern)	Zirzipanagrion quadriordinum, P. brunonis (114993)
Early/Lower Toarcian	183.0 - 182.0	Germany	Libellula sp. (123987) P. brunonis (114991)
Early/Lower Toarcian	183.0 - 182.0	Germany (Niedersachsen)	P. grasselensis (107321)
Falciferum	183.0 - 182.0	Luxembourg	Zirzipanagrion sp., P. bascharagensis (105427)
Toarcian	182.7 - 174.1	Germany	P. sp. 1, P. sp. 2, P. brunonis, P. pascholdi (104056)
Callovian - Oxfordian	166.1 - 157.3	Kazakhstan (Chimkent)	P. grandis (105971) P. handlirschi, P. angustovenosus (114692) P. karatauensis, P. kazakhstanensis (106101) P. kazakhstanensis, P. pumilio (105426) P. sp., P. grandis, P. kazakhstanensis (113439)
Tithonian	152.1 - 145.0	Mongolia	P. sp. (104748 121394)
Early/Lower Tithonian	150.8 - 145.0	Germany (Bayern)	P. jurassicus (123144)
Late/Upper Hauterivian	136.4 - 130.0	United Kingdom (England)	P. cretacicus (106417)