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Megasecoptera
Taxonomy
Megasecoptera was named by Brongniart (1885).
It was assigned to Insecta by Riek (1976); to Palaeoptera by Martynov (1932), Carpenter (1992); to Palaeodictyopterida by Grimaldi and Engel (2005); and to Megasecopteromorpha by Béthoux (2020).
It was assigned to Insecta by Riek (1976); to Palaeoptera by Martynov (1932), Carpenter (1992); to Palaeodictyopterida by Grimaldi and Engel (2005); and to Megasecopteromorpha by Béthoux (2020).
Subtaxa
Adiaphtharsia, Alectoneuridae, Anchineuridae, Ancopteridae, Archaeoscolex, Archaeoscolex corneus, Arcioneuridae, Aspidohymenidae, Aspidothoracidae, Bardohymenidae, Brodiidae, Caulopteridae, Corydaloididae, Dictyoneurellidae, Engisopteridae, Eumegasecoptera, Foririidae, Ischnoptilidae, Lameereites, Mischopteridae, Moravohymenidae, Piesbergpteridae, Protohymenidae, Raphidiopsis, Scytohymenidae, Sphecopteridae, Sphecorydaloididae, Vorkutiidae, Xenopteraidae
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1885 | Megasecoptera Brongniart p. 63 |
| 1889 | Archaeoscolex Matthew p. 59 |
| 1889 | Archaeoscolex corneus Matthew p. 59 figs. Pl 4, fig 11 |
| 1924 | Protohymenoptera Tillyard p. 112 |
| 1929 | Synsecoptera Martynov |
| 1932 | Megasecoptera Martynov |
| 1932 | Protohymenoptera Martynov p. 2 |
| 1933 | Protohymenoptera Carpenter |
| 1937 | Protohymenoptera Martynov p. 8 |
| 1976 | Megasecoptera Riek |
| 1984 | Archaeoscolex Durden p. 176 |
| 1984 | Archaeoscolex corneus Durden p. 176 |
| 1992 | Megasecoptera Carpenter p. 45 |
| 2005 | Megasecoptera Grimaldi and Engel |
| 2020 | Megasecoptera Béthoux |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Or. †Megasecoptera Brongniart 1885
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Fm. †Alectoneuridae Kukalová-Peck 1975
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Fm. †Anchineuridae Carpenter 1963
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Fm. †Ancopteridae Kukalová-Peck 1975
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G. †Agaeoleptoptera Beckemeyer and Engel 2011
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†Agaeoleptoptera uniotempla Beckemeyer and Engel 2011
Fm. †Arcioneuridae Kukalová-Peck 1975
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Fm. †Aspidohymenidae Martynov 1930
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G. †Aspidohymen Martynov 1930
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†Aspidohymen chopardi Zalessky 1937
†Aspidohymen extensus Martynov 1930
Fm. †Aspidothoracidae Handlirsch 1919
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G. †Aspidothorax Brongniart 1894
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†Aspidothorax hispanicus Santos et al. 2025
†Aspidothorax permianus Sinitshenkova and Aristov 2020
†Aspidothorax triangularis Brongniart 1894
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Invalid names: Aspidothorax maculatus Brongniart 1894 [synonym], Aspidothorax nanus Handlirsch 1919 [synonym]
Invalid names: Protocapnia Brongniart 1885 [nomen nudum]
Fm. †Bardohymenidae Zalessky 1937
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G. †Alexahymen Kukalová-Peck 1972
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†Alexahymen aestatis Brauckmann 1991
†Alexahymen maruska Kukalová-Peck 1972
G. †Sylvohymen Martynov 1940
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†Sylvohymen ingens Carpenter 1947
†Sylvohymen marginatus Pecharová et al. 2020
†Sylvohymen peckae Brauckmann 1988
†Sylvohymen robustus Martynov 1940
†Sylvohymen sibiricus Kukalová-Peck 1972
Fm. †Brodiidae Handlirsch 1906
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G. †Brodia Scudder 1881
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†Brodia jogginsensis Prokop et al. 2017
†Brodia priscotincta Scudder 1881
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Invalid names: Brodia fasciata Handlirsch 1919 [synonym], Brodia furcata Handlirsch 1919 [synonym], Brodia nebulosa Handlirsch 1919 [synonym], Brodia nympha Handlirsch 1919 [synonym], Brodia petiolata Handlirsch 1919 [synonym], Brodia pictipennis Handlirsch 1919 [synonym], Brodia scudderi Handlirsch 1919 [synonym]
Fm. †Caulopteridae Kukalová-Peck 1975
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Fm. †Corydaloididae Handlirsch 1906
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G. †Corydaloides Brongniart 1885
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†Corydaloides scudderi Brongniart 1885
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Invalid names: Corydaliodes gracilior Handlirsch 1919 [synonym], Corydaloides brongniarti Handlirsch 1919 [synonym], Corydaloides longipennis Handlirsch 1919 [synonym], Corydaloides simplex Brongniart 1894 [synonym]
Invalid names: Corydaliodes Handlirsch 1919 [synonym]
Fm. †Dictyoneurellidae Kukalová-Peck 1975
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Fm. †Engisopteridae Kukalová-Peck 1975
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G. †Engisoptera Kukalová-Peck 1975
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†Engisoptera ivanae Rýznar 2021
†Engisoptera simplices Kukalová-Peck 1975
Subor. †Eumegasecoptera
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Fm. †Aykhalidae Sinitshenkova 1993
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G. †Sinopalaeopteryx Pecharová et al. 2015
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†Sinopalaeopteryx olivieri Pecharová et al. 2015
†Sinopalaeopteryx splendens Pecharová et al. 2015
Fm. †Brodiopteridae Carpenter 1963
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G. †Brodioptera Copeland 1957
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†Brodioptera cumberlandensis Copeland 1957
†Brodioptera pintoi Brauckmann et al. 2003
†Brodioptera sinensis Pecharová et al. 2015
†Brodioptera stricklani Nelson and Tidwell 1987
Fm. †Ischnoptilidae Carpenter 1951
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Fm. †Mischopteridae Handlirsch 1906
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G. †Mischoptera Brongniart 1894
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†Mischoptera douglassi Carpenter and Richardson 1968
†Mischoptera nigra Brongniart 1885
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Invalid names: Mischoptera concolor Handlirsch 1919 [synonym], Mischoptera infumata Handlirsch 1919 [synonym], Mischoptera ocellata Handlirsch 1919 [synonym], Mischoptera woodwardi Brongniart 1894 [synonym]
Invalid names: Woodwardia Brongniart 1885 [replaced]
G. †Psilothorax Brongniart 1894
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†Psilothorax longicauda Brongniart 1894
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Invalid names: Psilothorax pictus Handlirsch 1919 [synonym]
Fm. †Moravohymenidae Kukalová-Peck 1972
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G. †Issadohymen Sinitshenkova and Aristov 2013
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†Issadohymen ponomarenkoi Sinitshenkova and Aristov 2013
Fm. †Piesbergpteridae Rosová et al. 2022
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Fm. †Protohymenidae Tillyard 1924
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G. †Arroyohymen Prokop and Kukalová-Peck 2017
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†Arroyohymen splendens Prokop and Kukalová-Peck 2017
G. †Protohymen Tillyard 1924
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†Protohymen angustipennis Martynov 1932
†Protohymen anomalus Zimmerman 1962
†Protohymen bifurcatus Zimmerman 1962
†Protohymen carpenteri Novokshonov 1995
†Protohymen concinna Zalessky 1956
†Protohymen curvatus Carpenter 1947
†Protohymen elongatus Carpenter 1930
†Protohymen largus Carpenter 1947
†Protohymen latus Carpenter 1947
†Protohymen novokshonovi Pecharová and Prokop 2018
†Protohymen permianus Tillyard 1924
†Protohymen pictus Zimmerman 1962
†Protohymen readi Carpenter 1933
†Protohymen shafferi Zimmerman 1962
†Protohymen tenuis Carpenter 1939
†Protohymen venustus Carpenter 1947
Invalid names: Pseudohymen Martynov 1932 [synonym], Pseudohymenopsis Zalessky 1956 [synonym]
Fm. †Scytohymenidae Martynov 1937
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G. †Karoohymen Riek 1976
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†Karoohymen delicatulus Riek 1976
†Karoohymen minutus van Dijk and Geertsema 1999
Fm. †Sphecopteridae Carpenter 1951
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G. †Cyclocelis Brongniart 1894
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†Cyclocelis chatini Brongniart 1894
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Invalid names: Sphecoptera brongniartiana Handlirsch 1919 [synonym], Sphecoptera fenestrata Handlirsch 1919 [synonym]
†Cyclocelis devexa Sinitshenkova 2019
†Cyclocelis obscura Brongniart 1894
†Cycloscelis elegantissima Meunier 1908
G. †Sphecoptera Brongniart 1894
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†Sphecoptera brongniarti Meunier 1908
†Sphecoptera elegantissima Meunier 1908
†Sphecoptera gracilis Brongniart 1894
†Sphecoptera minor Brongniart 1894
†Sphecoptera pulchra Brongniart 1894
Invalid names: Cyclocelis acuta Brongniart 1894 [nomen dubium]
Fm. †Sphecorydaloididae Pinto 1994
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Fm. †Vorkutiidae Rohdendorf 1947
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G. †Vorkutia Rohdendorf 1947
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†Vorkutia dimina Novokshonov 1998
†Vorkutia tshernovi Rohdendorf 1947
Fm. †Xenopteraidae Pinto 1986
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G. †Sinitshenkovae Nel et al. 2019
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†Sinitshenkovae gallica Nel et al. 2019
†Sinitshenkovae huenickeni Pinto and de Ornellas 1978
Invalid names: Archaeoscolex Matthew 1889 [nomen dubium], Archaeoscolex corneus Matthew 1889 [nomen dubium], Protohymenoptera Tillyard 1924 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range: base of the Kinderscoutian to the top of the Changhsingian or 321.80000 to 251.90200 Ma
Collections (80 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Bashkirian | Argentina (La Rioja) | Xenoptera riojaensis (108150) | |
| Kinderscoutian | USA (Utah) | Brodioptera stricklani (122484) | |
| Marsdenian | Germany (Ruhr) | Sylvohymen peckae (110939) Sylvohymen pintoi (152121) | |
| Westphalian A | Canada (New Brunswick) | Archaeoscolex corneus (114468) | |
| Westphalian A | Canada (Nova Scotia) | Brodia jogginsensis (187589) Brodioptera cumberlandensis (122485) | |
| Westphalian A | United Kingdom (England) | Brodia priscotincta (167683) | |
| Westphalian | USA (Alabama) | Agaeoleptoptera uniotempla (81525) | |
| Westphalian B | United Kingdom (Staffordshire) | Brodia furcata, Brodia nebulosa, Brodia pictipennis, Brodia nympha (34906) Brodia priscotincta (167682) Brodia priscotincta, Brodia scudderi (152262) | |
| Westphalian B | France (Nord-Pas-de-Calais) | Dictyoneurella perfecta (152792) | |
| Westphalian B | United Kingdom (England) | Brodia fasciata, Brodia nympha (125238) | |
| Westphalian B - Westphalian C | China (Ningxia) | Brodioptera sinensis, Sinopalaeopteryx splendens, Sinopalaeopteryx olivieri, Namuroptera minuta, Microhymen inconspicuus (139576) | |
| Westphalian C | United Kingdom (England) | Brodia priscotincta (127525) | |
| Westphalian C | United Kingdom (Staffordshire) | Brodia priscotincta (167681) | |
| Westphalian C - Westphalian D | France | Megasecoptera indet. (122504) | |
| Westphalian C - Westphalian D | Germany (Lower Saxony) | Aspidothorax tristrata (122514) | |
| Westphalian D | Germany (Lower Saxony) | Aspidothorax aestatis (172341) Aspidothorax tristrata (122515) Piesbergptera niesseae (224159) | |
| Westphalian D | USA (Illinois) | Adiaphtharsia ferrea (146857) Carbohymen testai (127721) Eubrodia dabasinskasi (112637 151848) Eubrodia dabasinskasi, Mischoptera douglassi (151852) Lameereites curvipennis (112630) Mischoptera douglassi (122538 161914) Mischoptera sp. (124839) | |
| Kasimovian | USA (Kansas) | Megasecoptera indet. (122507) | |
| Kasimovian - Gzhelian | Argentina (San Luis) | Megasecoptera indet., Philiasptilon huenickeni, Sphecorydaloides lucchesei (35057) | |
| Gzhelian | Russian Federation (Sakha) | Aykhal helenae (126060) | |
| Gzhelian | Spain (Leon) | Anchineura hispanica (105374) | |
| Gzhelian | Spain (León) | Aspidothorax hispanicus (239637) | |
| Gzhelian | Russian Federation | Siberiohymen asiaticus (124584) | |
| Gzhelian | France | Aspidothorax triangularis, Aspidothorax maculatus, Aspidothorax nanus, Woodwardia nigra, Psilothorax longicauda, Sphecoptera gracilis, Cyclocelis minor, Cyclocelis chatini, Sphecoptera brongniartiana, Cyclocelis obscura, Ischnoptilus elegans, Corydaloides scudderi, Corydaloides brongniarti, Corydaloides longipennis, Corydaloides simplex, Corydaliodes gracilior, Mischoptera woodwardi, Cyclocelis acuta, Sphecoptera pulchra, Mischoptera concolor, Mischoptera ocellata, Mischoptera infumata, Psilothorax pictus, Sphecoptera fenestrata (103635) Mischoptera nigra, Sphecoptera brongniarti, Foriria maculata, Cycloscelis elegantissima (121979) | |
| Asselian | USA (New Mexico) | Arroyohymen splendens (189778) | |
| Asselian | Czech Republic | Alexahymen maruska, Moravohymen vitreus, Cauloptera colorata, Arcioneura juveniles, Anconeura havlatai, Ancoptera sp., Ancoptera permiana, Engisoptera simplices, Alectoneura europaea (114186) | |
| Asselian | Czech Republic (Moravia) | Engisoptera ivanae (217298) | |
| Sakmarian | China (Shanxi) | Paleohymen shanxiensis, Paleohymen renmazhuangensis, Sinohymen gancaoshanensis (140933) | |
| Artinskian | USA (Kansas) | Protohymen permianus (111320) Protohymen permianus, Permohymen schucherti (111219) Protohymen permianus, Permohymen schucherti, Protohymen elongatus (111217) Protohymen permianus, Permohymen schucherti, Protohymen tenuis (114152) Protohymen readi, Protohymen permianus, Permohymen schucherti, Protohymen elongatus (114151) Protohymen readi, Protohymen permianus, Protohymen elongatus (111319) | |
| Artinskian | USA (Oklahoma) | Calohymen permianus, Protohymen curvatus, Protohymen venustus, Protohymen largus (104824) Protohymen bifurcatus, Protohymen readi (104826) Protohymen curvatus, Protohymen largus (104823) Protohymen curvatus, Protohymen latus (104822) Protohymen shafferi, Protohymen anomalus, Protohymen pictus (104827) Sylvohymen ingens (104889) | |
| Artinskian | China (Shanxi) | Sunohymen xishanensis (140458) | |
| Artinskian | USA (Texas) | Actinohymen russelli (111540) | |
| Kungurian | China (Shanxi) | Anthohymen hejinensis (140456) | |
| Kungurian | Russian Federation (Komi Republic) | Taigahymen borealis (201684) | |
| Kungurian | Russian Federation (Komi republic) | Cyclocelis devexa (201683) Vorkutia tshernovi (160744) | |
| Kungurian | Russian Federation | Pseudohymenopsis concinna, Bardohymen magnipennifer (161939) Pseudohymenopsis minor (102504) Scytohymen extremus, Fragmohymen submissus (114207) Scytohymenidae indet., Oceanoptera elenae (115214) Vorkutia dimina (107787) | |
| Kungurian | France (Languedoc) | Sinitshenkovae gallica (105045) | |
| Kungurian | Russian Federation (Kishertskij) | Pseudohymen angustipennis (111188) | |
| Kungurian | Russian Federation (Suksun) | Protohymen novokshonovi (122753) Pseudohymen sp., Pseudohymen sylvaensis, Sylvohymen robustus (116641) Sylvohymen marginatus (113380) Sylvohymen sibiricus, Pseudohymen carpenteri (108206) Tshekardohymen martynovi (124241) | |
| Kungurian - Early/Lower Kazanian | Russian Federation | Scytohymen extremus (116612) | |
| Roadian | Russian Federation (Perm) | Aspidothorax permianus (210714) | |
| Roadian | Russian Federation (Kazan) | Aspidohymen extensus (116803) | |
| Roadian | Russian Federation | Aspidohymen chopardi (122871) | |
| Roadian | Russian Federation (Arkangelsk) | Ivahymen constrictus (111187) | |
| Wuchiapingian | Russian Federation (Vologda) | Issadohymen ponomarenkoi (104754) | |
| Changhsingian | South Africa (Natal) | Karoohymen delicatulus, Karoohymen minutus (124428) |