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Amarina

Insecta - Coleoptera - Carabidae

Taxonomy
Amarina was named by Zimmerman (1832). It is extant.

It was assigned to Zabrini by Bouchard et al. (2011), Webster (2016).

Subtaxa
Amara

Synonymy list
YearName and author
1832Amarina Zimmerman
2011Amarina Bouchard et al. p. 145
2016Amarina Webster

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
RankNameAuthor
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderAdephaga
familyCarabidae()
subfamilyHarpalinae()
tribeZabrini()
subtribeAmarinaZimmerman 1832
subtribeAmarinaZimmerman 1832

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subtr. Amarina Zimmerman 1832
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G. Amara Bonelli 1810
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Subg. Amara (Amara) Bonelli 1810
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†Amara (Amara) subaenea Lomnicki 1894
Subg. Amara (Bradytus) Stephens 1827
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Subg. Amara (Celia) Zimmerman 1832
Subg. Amara (Curtonotus) Stephens 1827
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Curtonotus alpinus Paykull 1790
Curtonotus fodinae Mannerheim 1825
Subg. Amara (Paracelia) Bedel 1899
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Amara aenea De Geer 1774
Amara arcticola Poppius 1906
Amara aulica Panzer 1796
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Invalid names: Carabus ruficornis De Geer 1774 [replaced]
Amara bifrons Gyllenhal 1810
Amara brunnea Gyllenhal 1810
†Amara cockerelli Wickham 1912
Amara conflata Le Conte 1855
†Amara danae Scudder 1900
Amara erratica Duftschmid 1812
†Amara foersteri Handlirsch 1907
+
Invalid names: Amara sinuata Förster 1891 [replaced]
Amara insignis Dejean 1831
Amara municipalis Duftschmid 1812
†Amara occlusa Scudder 1900
†Amara paleomelas Scudder 1879
†Amara pinguicula Heer 1862
†Amara powellii Scudder 1900
†Amara primigenia Heer 1862
†Amara princeps Heer 1862
†Amara procera Förster 1891
Amara pseudobrunnea Lindroth 1968
†Amara revocata Scudder 1900
†Amara sterilis Scudder 1900
†Amara veterata Scudder 1900
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Ypresian to the top of the Rancholabrean or 56.00000 to 0.01400 Ma
Minimum age of oldest fossil (stem group age): 48.07 Ma

Collections (72 total)


Time interval Ma Country or state Original ID and collection number
Ypresian56.0 - 48.07Canada (British Columbia) Amara sp. (31677) Nebria paleomelas (122190)
Late/Upper Eocene37.71 - 33.9USA (Colorado) Amara danae, Amara sterilis (122093)
Chadronian37.0 - 33.9USA (Colorado) Amara cockerelli (114150) Amara cockerelli, Amara danae, Amara revocata, Amara powellii, Nebria occlusa, Amara sterilis, Amara veterata (106384) Amara danae (106379) Amara powellii (113749) Amara powellii, Amara veterata (124624)
Early/Lower Oligocene33.9 - 27.3France Amara sinuata, Amara procera (121473)
Sarmatian12.8 - 11.63Germany (Baden-Württemberg) Amara princeps, Amara primigenia, Amara pinguicula (117252)
Late/Upper Miocene11.63 - 5.333Russian Federation Amara sp., Amara quenseli (113713)
Messinian7.246 - 5.333USA (Alaska) Amara sp. (123352)
Gelasian2.58 - 1.8Greenland Amara glacialis (160160) Amara spp. (160144 160146 160151 160153) Amara spp., Amara brunnea (160143) Amara spp., Amara quenseli, Amara pseudobrunnea, Amara brunnea (160109)
Early/Lower Pleistocene2.58 - 0.774Russian Federation Amara sp., Curtonotus alpinus, Amara interstitialis (163069) Curtonotus alpinus (163089)
Chibanian0.774 - 0.129Russian Federation Amara glacialis (163096) Curtonotus alpinus (163090 163091 163092 163093 163094 163099 163100) Curtonotus alpinus, Amara glacialis (163098) Curtonotus alpinus, Curtonotus fodinae (163095 163097)
Middle Pleistocene0.774 - 0.129United Kingdom Amara aenea (168266 168267) Amara bifrons (168265)
Rancholabrean0.21 - 0.014USA (California) Amara insignis (150336)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation Amara glacialis, Curtonotus fodinae (173203) Curtonotus alpinus (153435 153436 153437 153438 153439 153440 153441 153442 153443 153444 153445 163101 163102 163103 163104 163108 163109 163110 173202 173210 173211 173212 173213 173214 173215) Curtonotus alpinus, Amara glacialis (153446 173208) Curtonotus alpinus, Amara glacialis, Curtonotus fodinae, Amara arcticola (173201) Curtonotus alpinus, Amara quenseli (173216)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Amara subaenea, Amara boryslavica (153476)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Worcestershire) Amara spp., Amara quenseli, Amara interstitialis, Amara municipalis, Amara erratica, Amara torrida (163070) Amara torrida (163075)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation (Astrakhan) Amara sp., Amara aenea (125794)
Late/Upper Pleistocene0.129 - 0.0117USA (Washington) Amara conflata (167662)
Tarantian0.129 - 0.0117USA (Pennsylvania) Amara sp. (137204)