PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Harpalus

Insecta - Coleoptera - Carabidae

Taxonomy

Harpalus was named by Latreille (1802). It is extant. It is the type genus of Harpalii, Harpalini, Harpalina.

It was assigned to Melanchlaeni by Latreille (1802); to Carabiques by Dejean (1821); to Harpali by LeConte (1861); to Carabidae by Scudder (1900), Wickham (1917), Cockerell (1921); to Harpalina by Bouchard et al. (2011); and to Harpalini by Tamutis et al. (2011).

Species lacking formal opinion data

H. amputatus, H. vittatus

Synonymy list

Year	Name and author
1802	Harpalus Latreille p. 92
1821	Harpalus Dejean p. 14
1861	Harpalus LeConte p. 33
1900	Harpalus Scudder p. 34
1917	Harpalus Wickham p. 464
1921	Harpalus Cockerell
2011	Harpalus Bouchard et al. p. 129
2011	Harpalus Tamutis et al.

Is something missing? Join the Paleobiology Database and enter the data

Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia

Rank	Name	Author
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Neuropteriformia
—	Coleopterida
order	Coleoptera
—	Mesocoleoptera
—	Metacoleoptera
suborder	Adephaga
family	Carabidae	()
subfamily	Harpalinae	()
tribe	Harpalini
subtribe	Harpalina
genus	Harpalus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Harpalus Latreille 1802

show all | hide all

†Harpalus abolitus von Heyden and von Heyden 1866

Harpalus amputatus Say 1830

†Harpalus bruckmanni Heer 1862

†Harpalus conditus Scudder 1900

†Harpalus constrictus Heer 1862

†Harpalus deletus Oustalet 1874

†Harpalus dilatatus Theobald 1937

†Harpalus excavatus Förster 1891

†Harpalus lheritieri Piton 1940

†Harpalus maceratus Wickham 1911

†Harpalus nero Oustalet 1874

†Harpalus nitens Heer 1870

†Harpalus nuperus Scudder 1900

†Harpalus obtusus Förster 1891

†Harpalus oustaleti Meunier 1915

†Harpalus pleistocenicus Lomnicki 1894

Harpalus pusillus Motschulsky 1850

†Harpalus redivivus Wickham 1917

†Harpalus sinis Heer 1847

†Harpalus stierlini Heer 1862

†Harpalus stygius Heer 1862

†Harpalus tabidus Heer 1847

†Harpalus tardigradus Heer 1862

†Harpalus ulomaeformis Wickham 1917

†Harpalus veterum Cockerell 1921

Harpalus vittatus Gelber 1833

†Harpalus whitfieldii Scudder 1900

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Selandian to the top of the Late/Upper Pleistocene or 61.66000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 59.24 Ma

Collections (37 total)

Time interval	Ma	Country or state	Original ID and collection number
Danian - Selandian	66.0 - 59.24	Svalbard and Jan Mayen	Carabites nitens (122011)
Selandian	61.66 - 59.24	France (Auvergne)	H. lheritieri (113685)
Bridgerian	51.1 - 45.9	USA (Colorado)	H. veterum (114681)
Late/Upper Eocene	37.71 - 33.9	USA (Colorado)	H. whitfieldii (122093)
Priabonian	37.71 - 33.9	France	H. oustaleti (113709)
Chadronian	37.0 - 33.9	USA (Colorado)	H. maceratus (106381) H. nuperus, H. whitfieldii (106384) H. ulomaeformis, H. redivivus (124624)
Early/Lower Oligocene	33.9 - 27.3	France	H. excavatus (121474) H. obtusus (121473)
MP 23	33.9 - 27.3	France	H. nero (122535)
Rupelian	33.9 - 27.3	Germany	H. oustaleti, H. offusus, H. dilatatus (122407)
Chattian	27.3 - 23.04	France (Provence-Alpes-Côte d'Azur)	H. nero (122488) H. oustaleti, H. deletus (122487)
Late/Upper Oligocene	27.3 - 23.04	France (Puy-de-Dome)	H. nero (155204 155205)
MP 30	27.3 - 23.04	Germany	H. abolitus (123945)
Sarmatian	12.8 - 11.63	Germany (Baden-Württemberg)	H. bruckmanni (122868) H. sinis (115305) H. stierlini, H. tardigradus, H. stygius, H. constrictus (117252)
Sarmatian	12.8 - 11.63	Croatia	H. tabidus (117250)
Middle Pleistocene	0.774 - 0.129	United Kingdom	H. sp. (168266)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Colorado)	H. sp. (169353)
Late/Upper Pleistocene	0.129 - 0.0117	Ukraine	H. pleistocenicus (153476)
Late/Upper Pleistocene	0.129 - 0.0117	Canada (Ontario)	H. conditus (152577)
Late/Upper Pleistocene	0.129 - 0.0117	Russian Federation	H. amputatus (173215) H. pusillus (153443 163104) H. pusillus, H. amputatus (173216) H. vittatus (153438 153439 153442) H. vittatus, H. amputatus (173201) H. vittatus, H. pusillus (153446)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom	H. sp. (152473)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Pennsylvania)	H. sp. (137207)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (Worcestershire)	H. sp. (163070)