PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Brachymetopus

Trilobita - Proetida - Brachymetopidae

Taxonomy

Brachymetopus was named by M'Coy (1847).

It was assigned to Brachymetopidae by Hahn et al. (1985) and Jell and Adrain (2002); to Proetida by Sepkoski (2002); and to Brachymetopinae by Owens and Hammann (1990) and Vanderlaan and Ebach (2015).

Species

B. (Acutimetopus), B. (Brachymetopus), B. (Conimetopus), B. drevermanni, B. edwardsi, B. gaoanensis, B. germanicus, B. jayae, B. maccoyi, B. nodosus, B. pseudometopina, B. rhinorhachis, B. spiniferus, B. spinosus, B. strzeleckii, B. woodlandi, B. woodwardii

Species lacking formal opinion data

B. bormanus, B. coignouides, B. edwardsi, B. indianwellsensis, B. ouralicus, B. spinosiformis, B. sucellus, B. vosegus

Synonyms

Loeipyge was named by Kobayashi and Hamada (1979).

It was synonymized subjectively with Brachymetopus by Owens (2003).

It was assigned to Brachymetopinae by Owens and Hammann (1990); to Brachymetopidae by Jell and Adrain (2002); and to Proetida by Sepkoski (2002).
Iriania was named by Archbold (1981).

It was replaced with Narinia by Archbold (1997); it was synonymized subjectively with Brachymetopus by Jell and Adrain (2002).

It was assigned to Brachymetopidae by Archbold (1981).
Brachymetopina was named by Reed (1903).

It was synonymized objectively with Brachymetopus by Jell and Adrain (2002).

Synonymy list

Year	Name and author
1847	Brachymetopus M'Coy p. 229
1903	Brachymetopina Reed
1979	Loeipyge Kobayashi and Hamada
1981	Iriania Archbold p. 36
1985	Brachymetopus Hahn et al.
1990	Brachymetopus Owens and Hammann
1990	Loeipyge Owens and Hammann
2002	Brachymetopus Jell and Adrain p. 351
2002	Loeipyge Jell and Adrain p. 399
2002	Brachymetopus Sepkoski
2002	Loeipyge Sepkoski
2015	Brachymetopus Vanderlaan and Ebach p. 12

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829

Rank	Name	Author
subphylum	Artiopoda	(Hou and Bergstrom)
class	Trilobita	Walch 1771
order	Proetida	Fortey and Owens 1975
superfamily	Aulacopleuroidea	(Angelin 1854)
family	Brachymetopidae	Prantl and Pribyl 1951
subfamily	Brachymetopinae	Prantl and Pribyl 1950
genus	Brachymetopus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Brachymetopus M'Coy 1847

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Subg. †Brachymetopus (Acutimetopus) Hahn and Hahn 1985

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†Brachymetopus (Acutimetopus) jesenicianus Hahn and Hahn 1977

Subg. †Brachymetopus (Brachymetopus) M'Coy 1847

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†Brachymetopus (Brachymetopus) gloriosus Engel and Morris 1996

†Brachymetopus (Brachymetopus) intermedius Engel and Morris 1992

†Brachymetopus (Brachymetopus) medinodulosus Engel and Morris 1992

Subg. †Brachymetopus (Conimetopus) Hahn and Hahn 1985

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†Brachymetopus (Conimetopus) alekseevi Mychko and Savchuk 2019

†Brachymetopus (Conimetopus) arcticus Hahn and Hahn 1992

†Brachymetopus (Conimetopus) ouralica de Verneuil 1845

†Brachymetopus drevermanni Hahn 1964

†Brachymetopus edwardsi Goldring 1958

†Brachymetopus gaoanensis Zhang 1982

†Brachymetopus germanicus Hahn 1964

†Brachymetopus jayae Archbold 1981

†Brachymetopus maccoyi Portlock 1843

†Brachymetopus maccoyi spinimarginatus Hahn and Hahn 1969

†Brachymetopus nodosus Wilson 1979

†Brachymetopus pseudometopina Gauri and Ramovs 1964

†Brachymetopus rhinorhachis Brezinski 1992

†Brachymetopus spiniferus Kobayashi and Hamada 1979

†Brachymetopus spinosus Herrick 1888

†Brachymetopus strzeleckii M'Coy 1847

†Brachymetopus woodlandi Owens 1984

†Brachymetopus woodwardii Whidborne 1896

Invalid names: Brachymetopina Reed 1903 [objective synonym], Iriania Archbold 1981 [synonym], Loeipyge Kobayashi and Hamada 1979 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

low Mg calcitec

Composition 2:

chitinc

Ontogeny:

molting, addition of partsc

Environment:

marinec

Locomotion:

fast-movingc

Life habit:

low-level epifaunalc

Diet:

deposit feedero

Vision:

well-developedf

Created:

2009-12-19 02:32:28

Modified:

2009-12-19 04:32:28

Source: f = family, o = order, c = class

References: Fortey and Owens 1999, Aberhan et al. 2004, Whittington et al. 1997

Age range: base of the Late/Upper Famennian to the top of the Wuchiapingian or 364.70000 to 254.17000 Ma

Collections (63 total)

Time interval	Ma	Country or state	Original ID and collection number
Famennian	372.2 - 358.9	USA (Pennsylvania)	B. gemma (54952)
Famennian	372.2 - 358.9	Germany (Nordrhein-Westfalen)	B. drevermanni (38296 38298 38299 38300 38302)
Late/Upper Famennian	364.7 - 360.7	Germany (Nordrhein-Westfalen)	B. drevermanni (38304)
Late/Upper Famennian	364.7 - 360.7	USA (Missouri)	B. spinosiformis (6029)
Late/Upper Famennian	364.7 - 360.7	China (Guangxi)	B. oblongus, B. germanicus, B. woodwardii (222338)
Hastarian	358.9 - 353.8	United Kingdom (England)	B. woodwardii (229660 229661 229662)
Hastarian	358.9 - 353.8	Germany (Nordrhein-Westfalen)	B. germanicus (38293)
Hastarian	358.9 - 353.8	France	B. germanicus (38642)
Hastarian	358.9 - 353.8	China (Guangxi)	B. germanicus (222332) B. maccoyi (222341)
Kinderhookian	358.9 - 352.0	USA (Ohio)	B. nodosus (229350)
Kinderhookian	358.9 - 352.0	USA (New Mexico)	B. indianwellsensis (64752)
Kinderhookian	358.9 - 352.0	USA (Missouri)	B. sp. (7390)
Tournaisian	358.9 - 346.7	Australia (New South Wales)	B. regularis (217053)
Tournaisian	358.9 - 346.7	Australia (Queensland)	B. intermedius (217040) B. regularis, B. dunstani, B. intermedius (223681)
Tournaisian - Visean	358.9 - 330.9	Australia (New South Wales)	B. ararat (217060) B. medinodulosus (217058)
Ivorian	353.8 - 346.7	Germany	B. ouralicus (37133) B. senckenbergianus, B. maccoyi, B. maccoyii (37132)
Ivorian	353.8 - 346.7	Belgium	B. maccoyi (37123) B. maccoyi, B. maccoyii (37127) B. ouralicus, B. sucellus, B. maccoyi (37122) B. sucellus, B. maccoyi (37124)
Ivorian	353.8 - 346.7	Spain (León)	B. maccoyi (67109)
Ivorian	353.8 - 346.7	Spain (Léon)	B. maccoyi (67110)
Ivorian - Chadian	353.8 - 340.0	Ireland	B. maccoyi (76501)
Ivorian - Holkerian	353.8 - 340.0	France	B. vosegus, B. maccoyi (66732)
Arundian - Holkerian	346.7 - 340.0	Ireland	B. sp. (76517)
Chadian	346.7 - 340.0	USA (Tennessee)	B. sp. (6914)
Chadian	346.7 - 340.0	United Kingdom (England)	B. maccoyi (5962 5963 5964)
Chadian - Arundian	346.7 - 340.0	USA (Tennessee)	B. sp. (6916)
Chadian - Arundian	346.7 - 340.0	USA (Kentucky)	B. sp. (562)
Visean	346.7 - 330.9	Belgium	B. ouralicus, B. maccoyi (37131)
Visean	346.7 - 330.9	Belgium (Antwerpen)	B. bormanus (38848)
Visean	346.7 - 330.9	Australia (New South Wales)	B. gloriosus (217075)
Visean	346.7 - 330.9	Germany	B. coignouides (66733)
Asbian	340.0 - 336.0	United Kingdom (England)	B. ornatus (5965) B. ornatus, B. parvus (5966) B. ouralicus (5995)
Asbian	340.0 - 336.0	Australia (New South Wales)	B. strzeleckii (133349)
Asbian	340.0 - 336.0	Belgium	B. bormanus (42593)
Asbian - Brigantian	340.0 - 330.9	United Kingdom (England)	B. ornatus (28383)
Bashkirian	323.2 - 315.2	Spain	B. sp. (162349)
Yeadonian	320.0 - 318.7	United Kingdom (England)	B. edwardsi (42757)
Gzhelian	303.7 - 298.9	USA (Alaska)	B. pseudometopina (100287)
Gzhelian	303.7 - 298.9	Slovenia	B. jesenicianus (67114) B. pseudometopina (67116) B. pseudometopina, B. jesenicianus (67112)
Asselian	298.9 - 295.5	Thailand	Loeipyge spinifer (133249)
Asselian - Sakmarian	298.9 - 290.1	Russian Federation (Bashkortostan)	B. alekseevi (75594)
Kungurian	279.3 - 272.3	USA (Texas)	B. rhinorhachis (75865)
Kungurian	279.3 - 272.3	Indonesia (Irian Jaya)	Iriania jayae (81375)
Roadian	272.3 - 268.8	Ukraine	B. sp. (119279)
Wuchiapingian	259.9 - 254.17	China (Anhui)	B. gaoanensis (102110)