PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Charonoscaphinae

Insecta - Coleoptera - Coptoclavidae

Taxonomy

Charonoscaphinae was named by Ponomarenko (1977). Its type is Charonoscapha.

It was assigned to Coptoclavidae by Ponomarenko (1977), Soriano et al. (2007), Bouchard et al. (2011) and Cai et al. (2022).

Subtaxa

Charonoscapha (type genus), Charonoscaphidia

Synonymy list

Year	Name and author
1977	Charonoscaphinae Ponomarenko p. 32
2007	Charanoscaphinae Soriano et al. p. 527
2011	Charonoscaphinae Bouchard et al.
2022	Charonoscaphinae Cai et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924

Rank	Name	Author
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Neuropteriformia
—	Coleopterida
order	Coleoptera
—	Mesocoleoptera
—	Metacoleoptera
suborder	Adephaga
family	Coptoclavidae	Ponomarenko 1961
subfamily	Charonoscaphinae
subfamily	Charonoscaphinae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Charonoscaphinae Ponomarenko 1977

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G. †Charonoscapha Ponomarenko 1977

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†Charonoscapha grossa Ponomarenko 1977

†Charonoscapha ovata Ponomarenko 1977

G. †Charonoscaphidia Ponomarenko 1977

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†Charonoscaphidia elongata Ponomarenko 1977

Diagnosis

Reference	Diagnosis
C. Soriano et al. 2007	Large to medium-sized beetles. Metacoxae without femoral plates. Metatibiae broadened and flattened, tarsi flattened but not broadened, narrower than tibiae. Metatibiae and tarsi with long natatory hairs.

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range: base of the Callovian to the top of the Oxfordian or 166.10000 to 157.30000 Ma

Collections (7 total)

Time interval	Ma	Country or state	Original ID and collection number
Callovian - Oxfordian	166.1 - 157.3	Kazakhstan (Chimkent)	Charonoscapha grossa (105970) Charonoscapha grossa, Charonoscapha ovata (105426) Charonoscapha ovata (113439 115717) Charonoscapha ovata, Charonoscaphidia elongata (105971) Charonoscaphidia elongata (106102 114238)