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Coptoclavinae

Insecta - Coleoptera - Coptoclavidae

Taxonomy
Coptoclavinae was named by Ponomarenko (1961). Its type is Coptoclava.

It was assigned to Coptoclavidae by Soriano et al. (2007), Zhao et al. (2018), Cai et al. (2022), Bao et al. (2025).

Synonymy list
YearName and author
1961Coptoclavinae Ponomarenko
2007Coptoclavinae Soriano et al. p. 529
2018Coptoclavinae Zhao et al. p. 207
2022Coptoclavinae Cai et al.
2025Coptoclavinae Bao et al. p. 403

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
RankNameAuthor
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderAdephaga
familyCoptoclavidaePonomarenko 1961
subfamilyCoptoclavinaePonomarenko 1961
subfamilyCoptoclavinaePonomarenko 1961

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Coptoclavinae Ponomarenko 1961
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G. †Bolbonectes Ponomarenko 1987
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†Bolbonectes intermedius Ponomarenko 1987
†Bolbonectes occidentalis Ponomarenko 1993
†Bolbonectus lithographicus Ponomarenko and Martínez-Delclòs 2000
G. †Coptoclava Ping 1928
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†Coptoclava longipoda Ping 1928
G. †Hoyaclava Soriano et al. 2007
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†Hoyaclava buscalionae Soriano et al. 2007
G. †Megacoptoclava Ponomarenko and Martínez-Delclòs 2000
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†Megacoptoclava longiurogomphia Ponomarenko and Martínez-Delclòs 2000
G. †Tigrivia Bao et al. 2023
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†Tigrivia baii Bao et al. 2023
Diagnosis
ReferenceDiagnosis
C. Soriano et al. 2007Large beetles. Metacoxae without femoral plates. Meso- and metatarsi flattened and broadened, not narrower than tibiae and without natatory hairs.
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range: base of the Oxfordian to the top of the Albian or 161.50000 to 100.50000 Ma

Collections (39 total)


Time interval Ma Country or state Original ID and collection number
Oxfordian161.5 - 154.8Russian Federation (Buryat) Bolbonectes occidentalis (105017)
Tithonian149.2 - 143.1Russian Federation Bolbonectes intermedius (123957 125408)
Hauterivian132.6 - 125.77Russian Federation (Chita) Coptoclava longipoda (152177)
Late/Upper Hauterivian132.6 - 125.77China (Hebei) Coptoclava longipoda (111077)
Barremian125.77 - 121.4Russian Federation (Chita) Bolbonectes intermedius (151642) Coptoclava longipoda (152180)
Early/Lower Barremian125.77 - 121.4Spain Bolbonectus lithographicus (123437)
Early/Lower Barremian125.77 - 121.4China Coptoclava longipoda (235409)
Late/Upper Barremian125.77 - 121.4China (Jilin) Coptoclava longipoda (155010)
Late/Upper Barremian125.77 - 121.4Spain (Castilla-La Mancha) Hoyaclava buscalionae, Megacoptoclava longiurogomphia (28006)
Early/Lower Aptian121.4 - 119.57Russian Federation (Chita) Coptoclava longipoda (152179)
Early/Lower Aptian121.4 - 119.57China (Zhejiang) Coptoclava longipoda (155611)
Aptian121.4 - 113.2China (Jilin) Coptoclava longipoda (151876 172391)
Aptian121.4 - 113.2China (Hebei) Tigrivia baii (241724)
Aptian121.4 - 113.2Mongolia Coptoclava longipoda (123933)
Aptian121.4 - 113.2Russian Federation (Chita) Coptoclava longipoda (126001 138148)
Aptian121.4 - 113.2Russian Federation Coptoclava longipoda (124811)
Aptian121.4 - 113.2China (Nei Mongol) Coptoclava longipoda (156008)
Aptian121.4 - 113.2Russian Federation (Buryatia) Coptoclava longipoda (105969 113350)
Aptian121.4 - 113.2China (Gansu) Coptoclava longipoda (154779)
Aptian121.4 - 113.2China (Liaoning) Coptoclava longipoda (150814 151536)
Aptian121.4 - 113.2China (Shandong) Coptoclava longipoda (106313 154887 154888 154890 154894 154897 156554 156555 156601 161254)
Albian113.2 - 100.5South Korea Coptoclava sp. (184781 184782 184783)