PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Paleoamblypygi

Arachnida - Amblypygi

Taxonomy

Paleoamblypygi was named by Weygoldt (1996). It is extant.

It was assigned to Amblypygi by Dunlop et al. (2013), Engel and Grimaldi (2014), Garwood et al. (2017), Dunlop (2018), Prendini (2025).

Subtaxa

Burmacharon, Graeophonus, Paracharontidae, Paracharontoidea, Weygoldtinidae

Synonyms

Libellula carbonaria was named by Scudder (1876). It is an impression. Its type locality is Cossett's Pit, Sydney coalfield, Cape Breton, which is in a Westphalian C/Westphalian D terrestrial shale in Canada. It is the type species of Graeophonus.

It was recombined as Graeophonus carbonarius by Scudder (1890), Petrunkevitch (1913), Petrunkevitch (1953), Dunlop et al. (2013); it was considered a nomen dubium by Scudder (1885), Dunlop (2018), Prendini (2025).
Graeophonus was named by Scudder (1890). Its type is Libellula carbonaria.

It was considered a nomen dubium by Dunlop (2018), Prendini (2025).

It was assigned to Paracharontidae by Dunlop et al. (2013), Engel and Grimaldi (2014); and to Paleoamblypygi by Garwood et al. (2017).
Paracharontoidea was named by Weygoldt (1996). It is extant.

It was considered an invalid subgroup of Paleoamblypygi by Garwood et al. (2017).

It was assigned to Paleoamblypygi by Engel and Grimaldi (2014).

Synonymy list

Year	Name and author
1876	Libellula carbonaria Scudder p. 89 fig. 2
1890	Graeophonus Scudder p. 453
1890	Graeophonus carbonarius Scudder p. 454 figs. Pl 40, figs 2,3,6
1913	Graeophonus carbonarius Petrunkevitch p. 67 figs. Pl 5, figs 27-29; text-figs 29-31
1953	Graeophonus carbonarius Petrunkevitch p. 99
1996	Paleoamblypygi Weygoldt
1996	Paracharontoidea Weygoldt
2013	Graeophonus Dunlop et al. p. 190
2013	Graeophonus carbonarius Dunlop et al. p. 190
2013	Paleoamblypygi Dunlop et al. p. 190
2014	Graeophonus Engel and Grimaldi p. 3
2014	Paleoamblypygi Engel and Grimaldi p. 3
2014	Paracharontoidea Engel and Grimaldi p. 3
2017	Paleoamblypygi Garwood et al. p. 11
2017	Graeophonus Garwood et al. p. 12
2018	Paleoamblypygi Dunlop p. 18
2025	Paleoamblypygi Prendini p. 397

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
subphylum	Chelicerata	Heymons 1901
—	Euchelicerata	Weygoldt and Paulus 1979
—	Prosomapoda

Rank	Name	Author
—	Planaterga
—	Dekatriata
—	Sclerophorata	Kamenz et al. 2011
class	Arachnida	Lamarck 1801
—	Pantetrapulmonata
subclass	Tetrapulmonata	Schultz 1990
—	Schizotarsata
—	Pedipalpi	Borner 1904
order	Amblypygi	Thorell 1883
suborder	Paleoamblypygi	Weygoldt 1996
suborder	Paleoamblypygi	Weygoldt 1996

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subor. Paleoamblypygi Weygoldt 1996

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G. †Burmacharon Hu et al. 2020

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†Burmacharon dunlopi Hu et al. 2020

Fm. †Weygoldtinidae Dunlop 2018

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G. †Weygoldtina Dunlop 2018

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†Weygoldtina anglica Pocock 1911

†Weygoldtina scudderi Pocock 1911

Invalid names: Graeophonus Scudder 1890 [nomen dubium], Graeophonus carbonarius Scudder 1876 [nomen dubium], Paracharontidae Weygoldt 1996 [empty], Paracharontoidea Weygoldt 1996 [invalid subgroup]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Grouping:

solitaryc

Environment:

terrestrialc

Locomotion:

actively mobilec

Created:

2017-04-17 09:55:58

Modified:

2017-04-17 09:55:58

Source: c = class

Reference: Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Westphalian A to the top of the Early/Lower Cenomanian or 318.70000 to 93.90000 Ma
Minimum age of oldest fossil (stem group age): 316.9 Ma

Collections (11 total)

Time interval	Ma	Country or state	Original ID and collection number
Westphalian A	318.7 - 316.9	Canada (Nova Scotia)	Graeophonus carbonarius (146858)
Westphalian B	316.9 - 314.6	United Kingdom (England)	Graeophonus anglicus (125238) Graeophonus sp. (127286)
Westphalian B	316.9 - 314.6	United Kingdom (Staffordshire)	Graeophonus anglicus (34906 34907 34908 34935)
Westphalian C	314.6 - 309.8	United Kingdom	Graeophonus anglicus (34909)
Westphalian C - Westphalian D	314.6 - 307.4	Canada (Nova Scotia)	Libellula carbonaria (122607)
Westphalian D	309.8 - 307.4	USA (Illinois)	Graeophonus carbonarius (21981)
Early/Lower Cenomanian	100.5 - 93.9	Myanmar	Burmacharon dunlopi (161726)