PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Cleiothyridina

Rhynchonellata - Athyridida - Athyrididae

Taxonomy

Cleiothyridina was named by Buckman (1906) [Sepkoski's age data: ]. Its type is Atrypa pectinifera. It is the type genus of Cleiothyridininae.

It was reranked as Athyris (Cleiothyridina) by Reed (1944).

It was assigned to Athyris by Reed (1944); to Athyridinae by Brunton and Champion (1974); to Spiriferida by Sepkoski (2002); to Cleiothyridininae by Williams et al. (2002); and to Athyrididae by Savage et al. (1978), Shen et al. (2003).

Species

Athyris (Cleiothyridina) bajtuganensis, Athyris (Cleiothyridina) epigona, Athyris (Cleiothyridina) warchensis, Cleiothyridellina accoliformis, C. accola, C. acutomarginalis, C. ailakensis, C. anabathra, C. attenuata, C. bajkurica, C. baracoodensis, C. barbata, C. capillata, C. circularis, C. ciriacksi, C. corculum, C. dalmiriensis, C. davidsoni, C. dilimensis, C. echidniformis, C. excavata, C. fimbriata, C. glenparkensis, C. globulina, C. grossula, C. hayasakai, C. hirsuta, C. interposita, C. intonsa, C. laminosa, C. laqueata, C. macleayana, C. maynci, C. milleri, C. mulsa, C. nana, C. nielseni, C. nikolaevi, C. obmaxima, C. orbicularis, C. ovalis, C. pectinifera, C. perthensis, C. pijaensis, C. pilularis, C. rara, C. rectimarginata, C. saraiensis, C. semiconcava, C. seriata, C. shenshuensis, C. simulans, C. solovjevae, C. subexpansa, C. sublamellosa, C. tenuilineata, C. tribulosa, C. tschironensis, C. uralica, C. xetriformis, C. zhexiensis, Spirifer deroissyi

Species lacking formal opinion data

C. angelicoides, C. coloradoensis, C. crassicardinalis, C. deroissii, C. elegans, C. glabristria, C. humerosa, C. incrassata, C. media, C. pecossi, C. segmentata, C. squamosa, C. transversa

Synonyms

Cleiothyridellina was named by Waterhouse (1978).

It was synonymized subjectively with Cleiothyridina by Williams et al. (2002).
Cleiothyris was named by Phillips (1841). Its type is Atrypa pectinifera.

It was replaced with Cleiothyridina by Brunton (1976).

Synonymy list

Year	Name and author
1841	Cleiothyris Phillips
1906	Cleiothyridina Buckman
1944	Athyris (Cleiothyridina) Reed p. 253
1974	Cleiothyridina Brunton and Champion p. 828
1978	Cleiothyridina Savage et al. p. 385
1978	Cleiothyridellina Waterhouse p. 82
2002	Cleiothyridina Sepkoski
2002	Cleiothyridina Williams et al. p. 1510
2003	Cleiothyridina Shen et al. p. 90

Is something missing? Join the Paleobiology Database and enter the data

Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
—	Lophophorata
—	Panbrachiopoda	Carlson and Cohen 2020
phylum	Brachiopoda	Cuvier 1805

Rank	Name	Author
subphylum	Rhynchonelliformea	Williams et al. 1996
class	Rhynchonellata	Williams et al. 1996
order	Athyridida	Boucot et al. 1964
suborder	Athyrididina	Boucot et al. 1964
superfamily	Athyridoidea	McCoy 1844
family	Athyrididae	McCoy 1841
subfamily	Cleiothyridininae
genus	Cleiothyridina	Buckman 1906

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Cleiothyridina Buckman 1906

show all | hide all

†Athyris (Cleiothyridina) bajtuganensis Netschajew 1911

†Athyris (Cleiothyridina) epigona Abich 1878

†Athyris (Cleiothyridina) semiconcava dokriensis Reed 1944

†Athyris (Cleiothyridina) warchensis Reed 1944

†Cleiothyridina accola Reed 1944

†Cleiothyridina acutomarginalis Waagen 1883

†Cleiothyridina ailakensis Reed 1925

†Cleiothyridina anabathra Waterhouse 1968

†Cleiothyridina attenuata Cooper 1957

†Cleiothyridina bajkurica Chernyak 1963

†Cleiothyridina baracoodensis Etheridge Jr. 1903

†Cleiothyridina barbata Chronic 1949

†Cleiothyridina capillata Waagen 1883

†Cleiothyridina circularis Ustritsky 1960

†Cleiothyridina ciriacksi Cooper and Grant 1976

†Cleiothyridina corculum Lee and Su 1980

†Cleiothyridina dalmiriensis Reed 1944

†Cleiothyridina davidsoni Rigaux 1873

†Cleiothyridina dilimensis Grunt 1977

†Cleiothyridina echidniformis Waterhouse 1983

†Cleiothyridina excavata Grabau 1931

†Cleiothyridina fimbriata Phillips 1836

†Cleiothyridina glenparkensis Weller 1914

†Cleiothyridina globulina Waagen 1883

†Cleiothyridina grossula Waagen 1883

†Cleiothyridina hayasakai Tazawa 2010

†Cleiothyridina hirsuta Hall 1857

†Cleiothyridina interposita Reed 1944

†Cleiothyridina intonsa Chronic 1949

†Cleiothyridina laminosa Fang 1994

†Cleiothyridina laqueata Waterhouse 1968

†Cleiothyridina macleayana Etheridge Jr. 1889

†Cleiothyridina maynci Dunbar 1955

†Cleiothyridina milleri Savage et al. 1978

†Cleiothyridina mulsa Cooper and Grant 1976

†Cleiothyridina nana Cooper and Grant 1976

†Cleiothyridina nielseni Dunbar 1955

†Cleiothyridina nikolaevi Grunt 1977

†Cleiothyridina obmaxima McChesney 1861

†Cleiothyridina orbicularis McChesney 1859

†Cleiothyridina ovalis Shi and Archbold 1993

†Cleiothyridina pectinifera Sowerby 1841

†Cleiothyridina perthensis Archbold 1997

†Cleiothyridina pijaensis Waterhouse 1983

†Cleiothyridina pilularis Cooper and Grant 1976

†Cleiothyridina rara Cooper and Grant 1976

†Cleiothyridina rectimarginata Cooper and Grant 1976

†Cleiothyridina saraiensis Reed 1944

†Cleiothyridina semiconcava Waagen 1883

†Cleiothyridina seriata Grant 1976

†Cleiothyridina shenshuensis Lee and Su 1980

†Cleiothyridina simulans Reed 1931

†Cleiothyridina solovjevae Grunt 1977

†Cleiothyridina subexpansa Waagen 1883

†Cleiothyridina sublamellosa Hall 1858

†Cleiothyridina tenuilineata Rowley 1900

†Cleiothyridina tribulosa Grant 1976

†Cleiothyridina tschironensis Kotlyar 1968

†Cleiothyridina uralica Grabau 1934

†Cleiothyridina xetriformis Reed 1944

†Cleiothyridina zhexiensis Liang 1982

†Spirifer deroissyi Leveille 1835

Invalid names: Cleiothyridellina Waterhouse 1978 [synonym], Cleiothyris Phillips 1841 [replaced]

Diagnosis

No diagnoses are available

Measurements

part	N	mean	minimum	maximum	median	s.d.
shell height	245	7.87	1.20	27.5	8.39	5.88
valve height	3	6.94	4.19	9.30	8.60	2.77

Composition:

low Mg calcitep

Environment:

marineuc

Locomotion:

stationarysubo

Attached:

yesp

Life habit:

low-level epifaunalsubo

Diet:

suspension feedersubo

Vision:

blindc

Created:

2018-11-10 18:06:04

Modified:

2018-11-10 18:06:04

Source: subo = suborder, c = class, p = phylum, uc = unranked clade

References: Nesnidal et al. 2013, Aberhan et al. 2004

Age range: base of the Frasnian to the top of the Changhsingian or 382.31000 to 251.90200 Ma

Collections (1271 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Late/Upper Givetian - Early/Lower Frasnian	383.4 - 379.0	Spain (Leon)	C. sp. (26894)
Frasnian - Famennian	382.31 - 358.86	Afghanistan (Vardak)	C. sp. (84635)
Late/Upper Frasnian	375.2 - 372.15	Belgium	C. davidsoni (92831)
Late/Upper Frasnian	375.2 - 372.15	USA (Nevada)	C. angelicoides (156526)
Early/Lower Famennian	372.15 - 368.5	Germany (Rhenish Massif)	C. deroissii, C. sp. (5960)
Early/Lower Famennian	372.15 - 368.5	China	C. submabrancea (162317)
Early/Lower Famennian	372.15 - 368.5	Belgium	C. davidsoni (92832)
Early/Lower Famennian	372.15 - 368.5	Canada (Alberta)	C. sp. (35541)
Famennian	372.15 - 358.86	China (Xinjiang)	C. sp. (3560)
Famennian	372.15 - 358.86	USA (Alaska)	C. milleri (40582 40583 40584)
Youngest occurrences
Time interval	Ma	Country or state	PBDB collection number
Wuchiapingian - Changhsingian	259.51 - 251.902	Pakistan	Athyris capillata, Athyris xetriformis, Athyris dalmiriensis, Athyris accola (127019) Athyris dalmiriensis (126751) Athyris dalmiriensis, Athyris gerardi (126759) Athyris gerardi (126757) Athyris globulina (126705 126709) Athyris grossula, Athyris pectinifera (126755) Athyris pectinifera (126752) Athyris subexpansa, Athyris capillata, Athyris interposita (127023) Athyris subexpansa, Athyris dalmiriensis (126746 126998) Athyris subexpansa, Athyris dalmiriensis, Athyris globulina (126760) Athyris subexpansa, Athyris dalmiriensis, Athyris interposita, Athyris accola (126986) Athyris subexpansa, Athyris dalmiriensis, Athyris pectinifera (126754) Athyris subexpansa, Athyris globulina (126747) Athyris subexpansa, Athyris grossula, Athyris capillata, Athyris dalmiriensis, Athyris globulina, Athyris pectinifera (126740) Athyris subexpansa, Athyris grossula, Athyris capillata, Athyris warchensis, Athyris pectinifera (126886) Athyris subexpansa, Athyris grossula, Athyris dalmiriensis, Athyris globulina (126750) Athyris subexpansa, Athyris grossula, Athyris dalmiriensis, Athyris globulina, Athyris gerardi (126753) Athyris subexpansa, Athyris grossula, Athyris dalmiriensis, Athyris simulans, Athyris globulina, Athyris gerardi (126762) Athyris subexpansa, Athyris grossula, Athyris xetriformis, Athyris dalmiriensis (126704) Athyris subexpansa, Athyris grossula, Athyris xetriformis, Athyris dalmiriensis, Athyris pectinifera, Athyris gerardi (127014) Athyris subexpansa, Athyris pectinifera (127017)
Changhsingian	254.14 - 251.902	China (Guangxi)	C. sp. 2 (95336 95338)
Changhsingian	254.14 - 251.902	China (Guizhou)	C. sp. 1 (136834)
Changhsingian	254.14 - 251.902	China (Xizang)	C. capillata, C. royssii (121595) C. deroyssii (51398 51401) C. ovalis (73602) C. ovalis, C. subexpansa (73600) C. sp. (117541) C. sp., C. ovalis (73601) C. subexpansa (73603 73605)
Changhsingian	254.14 - 251.902	Pakistan (Northwest Frontier)	C. capillata (41557 41558)
Changhsingian	254.14 - 251.902	China (Chongqing)	Athyris excavata (94984) C. sp. (149259 149281)
Changhsingian	254.14 - 251.902	New Zealand	C. laqueata (98149)
Changhsingian	254.14 - 251.902	Nepal	Cleiothyridellina accoliformis (43166 43167 43176 43177 43180 43184 43185 43188 43189 43191 43197 43198 43199 43216 43217 43220 43221 43227 43255 62441 62445 62447) C. pijaensis (120509 120520 120521 120523) C. pijaensis, C. echidniformis (120508 120517 120519 120522) C. sp. (43155 43243 43261 43276 43282 43285) C. sp. A, Cleiothyridellina accoliformis (43187) C. sp. B (43194) C. sp. C (43182) C. sp. D, Cleiothyridellina accoliformis (43170) C. subexpansa (99791 120443) C. subexpansa, Cleiothyridellina accoliformis (62444 120386)
Changhsingian	254.14 - 251.902	China (Hunan)	C. sp. 1 (162509)
Changhsingian	254.14 - 251.902	Pakistan	Athyris accola (126887) Athyris capillata (83145 90251 126883) Athyris dalmiriensis (126933) Athyris gerardi (126996) Athyris globulina (126989) Athyris grossula (126934 126981) Athyris grossula, Athyris pectinifera (126896) Athyris pectinifera (119657) Athyris pectinifera, Athyris royssii (89064 119608) Athyris subexpansa (81504 120160) Athyris subexpansa, Athyris accola, Athyris pectinifera, Athyris gerardi (126882) Athyris subexpansa, Athyris capillata (119637) Athyris subexpansa, Athyris globulina (126897) C. capillata (132761 132762)