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Protoleninae

Artiopoda - Redlichiida - Ellipsocephalidae

Taxonomy
Protolenidae was named by Richter and Richter. Its type is Protolenus.

It was considered an invalid subgroup of Ellipsocephalidae by Adrain (2011); it was reranked as the subfamily Protoleninae by Cederström et al. (2022), El Albani et al. (2024).

It was assigned to Redlichiida by Palmer and Gatehouse (1972); and to Ellipsocephalidae by Cederström et al. (2022), El Albani et al. (2024).

Synonymy list
p. 15
YearName and author
1972Protolenidae Palmer and Gatehouse p. 15
2022Protoleninae Cederström et al. p. 119
2024Protoleninae El Albani et al. p. S7

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
EcdysozoaAguinaldo et al. 1997
Panarthropoda
phylumArthropodaGravenhorst 1843
classArtiopodaHou and Bergstrom
RankNameAuthor
classTrilobitaWalch 1771
orderRedlichiidaRichter 1932
suborderRedlichiinaRichter 1932
superfamilyEllipsocephaloideaLin 1990
familyEllipsocephalidaeMatthew 1887
subfamilyProtoleninae(Richter and Richter)
subfamilyProtoleninae(Richter and Richter)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Protoleninae Richter and Richter
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G. †Anaxarete Geyer 2016
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Anaxarete senior Cobbold 1910
G. †Chorbusulina Lazarenko 1962
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Chorbusulina bella Lazarenko 1964
Chorbusulina wilkesi Palmer and Gatehouse 1972
G. †Cuneoaxiella Cederström et al. 2022
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Cuneoaxiella grandis Ahlberg and Bergström 1978
G. †Protolenus Matthew 1892
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Protolenus elegans Matthew 1892
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Invalid names: Protolenus paradoxoides Matthew 1892 [synonym]
Protolenus pustulatus Cobbold 1910
Invalid names: Bergeronia Matthew 1895 [objective synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitec
Composition 2: chitinc
Ontogeny: molting, addition of partsc
Environment: marinec
Locomotion: fast-movingo
Life habit: low-level epifaunalo
Diet: deposit feedero
Vision: well-developedf
Created: 2009-12-19 04:00:08
Modified: 2009-12-19 06:00:08
Source: f = family, o = order, c = class
References: Whittington et al. 1997, Hendy et al. 2009, Aberhan et al. 2004

Age range: base of the Branchian to the top of the Delamaran or 516.20000 to 505.30000 Ma

Collections (45 total)


Time interval Ma Country or state Original ID and collection number
Caerfai538.8 - 506.5Canada (New Brunswick) Protolenus articephalus, Protolenus elegans (34567 34751) Protolenus elegans (34569 34570 34571 34595 34596 34752)
Early/Lower Cambrian538.8 - 506.5Antarctica Chorbusulina wilkesi (154030)
Early/Lower Cambrian538.8 - 506.5Spain (Aragon) Protolenus pisidianus, Protolenus jilocanus (120550 120552)
Atdabanian521.0 - 516.0Russian Federation (Siberia) Protolenidae indet. (232892)
Atdabanian521.0 - 516.0Spain Protolenus schneideri (225087)
Atdabanian521.0 - 516.0Poland Protolenus bodzanti, Protolenus medius, Protolenus czarnockii (72261) Protolenus sp. (72268)
Atdabanian521.0 - 516.0Sweden Proampyx grandis (87121 87212)
Branchian516.2 - 506.5United Kingdom (England) Protolenus indet., Protolenus pustulatus (79928)
Lenian516.0 - 508.2Morocco Protolenus densigranulatus (161460) Protolenus dimarginatus, Protolenus hupei, Protolenus termierelloides (9801)
Stage 4514.5 - 506.5Morocco Protolenus sp. (235700)
Stage 4514.5 - 506.5Sweden Cuneoaxiella grandis (236571 236572 236573 236574 240025 240026 240027 240028 240029)
Toyonian512.1 - 508.2Spain (Zaragoza) Protolenus dimarginatus, Protolenus termierelloides, Protolenus jilocanus (233072)
Toyonian512.1 - 508.2Morocco Protolenus densigranulatus (9123)
Delamaran509.7 - 505.3Morocco Protolenus densigranulatus (174208 174210 174217 174219 174223 174291)
Amgan508.2 - 504.5Germany Protolenus bergstroemi (162993 162994 162996 162997)
Amgan508.2 - 504.5Poland Protolenus polonicus (72270)
Middle Cambrian506.5 - 497.0Poland Protolenus polonicus (72269)
Middle Cambrian506.5 - 497.0Italy (Iglesiente) Protolenus pisidianus (139878)