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Protoleninae

Trilobita - Redlichiida - Ellipsocephalidae

Taxonomy
Protolenidae was named by Richter and Richter. Its type is Protolenus.

It was considered an invalid subgroup of Ellipsocephalidae by Adrain (2011); it was reranked as the subfamily Protoleninae by Cederstrom et al. (2022) and El Albani et al. (2024).

It was assigned to Redlichiida by Palmer and Gatehouse (1972); and to Ellipsocephalidae by Cederstrom et al. (2022) and El Albani et al. (2024).

Synonymy list
p. 15
YearName and author
1972Protolenidae Palmer and Gatehouse p. 15
2022Protoleninae Cederstrom et al. p. 119
2024Protoleninae El Albani et al. p. S7

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
subphylumArtiopoda(Hou and Bergstrom)
RankNameAuthor
classTrilobitaWalch 1771
orderRedlichiidaRichter 1932
suborderRedlichiinaRichter 1932
superfamilyEllipsocephaloideaLin 1990
familyEllipsocephalidaeMatthew 1887
subfamilyProtoleninae(Richter and Richter)
subfamilyProtoleninae(Richter and Richter)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Protoleninae Richter and Richter
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G. †Anaxarete Geyer 2016
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Anaxarete senior Cobbold 1910
G. †Chorbusulina Lazarenko 1962
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Chorbusulina bella Lazarenko 1964
Chorbusulina subdita Palmer and Gatehouse 1972
Chorbusulina wilkesi Palmer and Gatehouse 1972
G. †Cuneoaxiella Cederstrom et al. 2022
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Cuneoaxiella grandis Ahlberg and Bergstrom 1978
G. †Protolenus Matthew 1892
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Subg. †Protolenus (Protolenus) Matthews 1892
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Protolenus pustulatus Cobbold 1910
Invalid names: Bergeronia Matthew 1895 [objective synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitec
Composition 2: chitinc
Ontogeny: molting, addition of partsc
Environment: marinec
Locomotion: fast-movingo
Life habit: low-level epifaunalo
Diet: deposit feedero
Vision: well-developedf
Created: 2009-12-19 04:00:08
Modified: 2009-12-19 06:00:08
Source: f = family, o = order, c = class
References: Hendy et al. 2009, Whittington et al. 1997, Aberhan et al. 2004

Age range: base of the Branchian to the top of the Delamaran or 517.10000 to 506.50000 Ma

Collections (40 total)


Time interval Ma Country or state Original ID and collection number
Caerfai538.8 - 509.0Canada (New Brunswick) Protolenus articephalus, Protolenus elegans (34567 34751) Protolenus elegans (34569 34570 34571 34595 34596 34752)
Early/Lower Cambrian538.8 - 509.0Spain (Aragon) Protolenus jilocanus, Protolenus pisidianus (120550 120552)
Early/Lower Cambrian538.8 - 509.0Antarctica Chorbusulina wilkesi, Chorbusulina subdita (154030)
Atdabanian521.0 - 516.0Russian Federation (Siberia) Protolenidae indet. (232892)
Atdabanian521.0 - 516.0Spain Protolenus schneideri (225087)
Atdabanian521.0 - 516.0Poland Protolenus bodzanti, Protolenus medius, Protolenus czarnockii (72261) Protolenus sp. (72268)
Atdabanian521.0 - 516.0Sweden Proampyx grandis (87121 87212)
Branchian517.1 - 509.0United Kingdom (England) Protolenus indet., Protolenus pustulatus (79928)
Lenian516.0 - 510.2Morocco Protolenus densigranulatus (161460) Protolenus dimarginatus, Protolenus hupei, Protolenus termierelloides (9801)
Stage 4514.0 - 509.0Sweden Cuneoaxiella grandis (236571 236572 236573 236574)
Stage 4514.0 - 509.0Morocco Protolenus sp. (235700)
Toyonian512.9 - 510.2Morocco Protolenus densigranulatus (9123)
Toyonian512.9 - 510.2Spain (Zaragoza) Protolenus jilocanus, Protolenus dimarginatus, Protolenus termierelloides (233072)
Delamaran511.2 - 506.5Morocco Protolenus densigranulatus (174208 174210 174217 174219 174223 174291)
Amgan510.2 - 504.5Poland Protolenus polonicus (72270)
Amgan510.2 - 504.5Germany Protolenus bergstroemi (162993 162994 162996 162997)
Middle Cambrian509.0 - 497.0Italy (Iglesiente) Protolenus pisidianus (139878)
Middle Cambrian509.0 - 497.0Poland Protolenus polonicus (72269)