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Ticholeptinae

Osteichthyes - Merycoidodontidae

Synonymy list
YearName and author
1941Ticholeptinae Schultz and Falkenbach p. 4
1996Ticholeptinae Stevens and Stevens p. 510
2007Ticholeptinae Stevens and Stevens p. 163
2009Ticholeptinae Morgan et al.
2011Ticholeptinae Lander and Lindsay p. 221
2023Ticholeptinae Stevens et al. p. 88

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
RankNameAuthor
Mammaliamorpha
Mammaliaformes
classMammalia
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Euungulata
Artiodactylamorpha
Artiodactyla()
suborderTylopoda()
familyMerycoidodontidae
subfamilyTicholeptinae
subfamilyTicholeptinae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Ticholeptinae Schultz and Falkenbach 1941
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G. †Paroreodon Thorpe 1921
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Paroreodon parvus Thorpe 1921
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Invalid names: Oreodontoides (Paroreodon) stocki Schultz and Falkenbach 1947 [synonym], Oreodontoides stocki Schultz and Falkenbach 1947 [synonym], Paroreodon marshi Thorpe 1921 [synonym]
Invalid names: Epigenetochoerus Schultz and Falkenbach 1968 [synonym]
G. †Phenacocoelus Peterson 1907
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Phenacocoelus kayi Schultz and Falkenbach 1950
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Invalid names: Hypsiops luskensis Schultz and Falkenbach 1950 [synonym]
Phenacocoelus typus Peterson 1907
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Invalid names: Phenacocoelus munroensis Peterson 1928 [synonym]
G. †Ticholeptus Cope 1878
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Ticholeptus bluei Schultz and Falkenbach 1947
Ticholeptus calimontanus Dougherty 1940
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Invalid names: Ticholeptus tooheyi Schultz and Falkenbach 1941 [synonym]
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Invalid names: Merychyus smithi Douglass 1903 [synonym], Merycochoerus obliquidens Cope 1886 [synonym], Poatrephes paludicola Douglass 1903 [synonym], Ticholeptus hypsodus Loomis 1924 [synonym], Ticholeptus hypsodus leadorensis Schultz and Falkenbach 1968 [synonym], Ticholeptus rileyi Schultz and Falkenbach 1941 [synonym], Ustatochoerus schrammi Schultz and Falkenbach 1941 [synonym]
Invalid names: Poatrephes Douglass 1903 [synonym]
Diagnosis
ReferenceDiagnosis
C. B. Schultz and C. H. Falkenbach 1941Medium to large size forms; skulls brachycephalic; supraoccipital wings incorporated in fan-shaped occipital region; tendency for a slight retraction of nasals (not to the same degree as in the Merycochoerinre); tympanic bullre large and flattened; teeth large and moder- ately hypsodont; premolars complicated.
M. S. Stevens et al. 2023The more advanced species are medium-sized oreodonts with brachycephalic skulls showing a fan-shaped occipital region, well-inflated auditory bullae, mesodont cheek teeth, and molarized premolars. The malar is prominent anterior to the orbit, causing a broad facial depression in the lacrimal region merging with the facial vacuity. The zygomatic arch is lightly constructed, and the nasals are moderately long with a slight retraction. The infraorbital foramen is above P3- P4. The rostrum is square and broad, not long and narrow as in the Ustatochoerinae. The postglenoid processes are larger and more robust than in the Ustatochoerinae. The premolars are elongated anteroposteriorly and are lower crowned than Merychyus, Mediochoerus, and Ustatochoerus. There is a small cusp on the posterolingual side of P2, the m1 is small, and m3 has a pronounced heel. Ticholeptus differs from Ustatochoerinae (formerly placed in the Ticholeptinae by Schultz and Falkenbach, 1941) in having lower crowned teeth, less reduction and molarization of premolars, which are also transversely broader, and only slight nasal retraction. The most primitive species (T. bluei, Paroreodon parvus, Phenacocoelus typus) do not show all these features, but do show the infraorbital foramen over the P3- P4, the deep narial notch and short nasals, and the large robust postglenoid processes.
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialo
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: herbivoref
Reproduction: viviparoussubc
Created: 2005-08-31 14:37:21
Modified: 2005-08-31 16:37:21
Source: f = family, o = order, subc = subclass, c = class, subp = subphylum
References: Hendy et al. 2009, Nowak 1999, Lander 1998, Carroll 1988, Lillegraven 1979

Age range: base of the Monroecreekian to the top of the Early/Lower Miocene or 24.70000 to 15.98000 Ma

Collections (71 total)


Time interval Ma Country or state Original ID and collection number
Arikareean29.5 - 18.5USA (Oregon) Paroreodon parvus (93080)
early Late Arikareean29.5 - 18.5USA (Oregon) Paroreodon sp. (36761 36763 36764 36766 36852 36854 36855 36856 170306)
Chattian27.3 - 23.04USA (Oregon) Paroreodon parvus (93079) Paroreodon sp. (17603)
Monroecreekian24.7 - 23.1USA (Nebraska) Phenacocoelus typus (17565)
Monroecreekian - Harrisonian24.7 - 18.5USA (Oregon) Paroreodon parvus (17528)
Harrisonian23.1 - 18.5USA (Wyoming) Hypsiops luskensis (17780) Phenacocoelus kayi (17840) Phenacocoelus typus (17794)
Harrisonian23.1 - 18.5USA (Florida) Phenacocoelus luskensis (18558)
Harrisonian23.1 - 18.5USA (Nebraska) Phenacocoelus sp. (17819)
Harrisonian - Hemingfordian23.1 - 16.3USA (Montana) Ticholeptus sp. (18811)
Early/Lower Miocene23.04 - 15.98USA (Oregon) Oreodontoides stocki (157001)
Miocene23.04 - 5.333USA (Oregon) Ticholeptus sp. (18898)
Miocene23.04 - 5.333USA (Nevada) Ticholeptus zygomaticus (18922 18925 18946)
Miocene23.04 - 5.333USA (California) Merychyus calimontanus (19477)
Early/Lower Hemingfordian18.5 - 16.3USA (Nebraska) Merychyus elegans (17978) Ticholeptus bluei (17935)
Hemingfordian18.5 - 16.3USA (California) Ticholeptus calimontanus (19367 19600)
Hemingfordian18.5 - 16.3USA (Nevada) Ticholeptus sp. (18991) Ticholeptus zygomaticus (18960 18961 18962 18963 18964 18965)
Hemingfordian18.5 - 16.3USA (Colorado) Ticholeptus sp. (17908)
Hemingfordian18.5 - 16.3USA (Nebraska) Ticholeptus hypsodus (17979 17999)
Barstovian16.3 - 12.5USA (Oregon) Merycoidodontidae indet. (18809) Ticholeptus obliquidens (19017 19019) Ticholeptus sp. (19056 19092) Ticholeptus zygomaticus (19027 19031)
Barstovian16.3 - 12.5USA (Nebraska) Ticholeptus sp. (18261 18284) Ustatochoerus schrammi (18124)
Barstovian16.3 - 12.5USA (California) Merychyus relictus (19513)
Barstovian16.3 - 12.5USA (Montana) Ticholeptus zygomaticus, Poatrephes paludicola (18894)
Barstovian16.3 - 12.5USA (Nevada) Ticholeptus sp. (19103)
Barstovian16.3 - 12.5USA (Idaho) Ticholeptus hypsodus (19013)
Barstovian16.3 - 12.5Canada (Saskatchewan) Ticholeptus sp. (18758 18759 18794 18797 18799)
Barstovian16.3 - 12.5USA (Texas) Ticholeptus rileyi (18619)
Early/Lower Barstovian16.3 - 12.5USA (Nebraska) Ticholeptus hypsodus (17904 17905 17955 17956 17972 17976 18004 18011 18020)
Early/Lower Barstovian16.3 - 12.5USA (Florida) Ticholeptus hypsodus (18549)
Early/Lower Barstovian16.3 - 12.5USA (Montana) Ticholeptus sp. (18803)