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Bembidiini

Insecta - Coleoptera - Carabidae

Taxonomy
Bembidiini was named by Stephens (1827). It is extant.

It was corrected as Bembidiides by Motschulsky (1864); it was reranked as the subfamily Bembidiinae by Hayashi et al. (2004).

It was assigned to Harpalidae by LeConte (1861); to Carabiques by Motschulsky (1864); to Carabidae by Hayashi et al. (2004); to Trechinae by Tamutis et al. (2011), Bouchard et al. (2011), Ortuño and Arillo (2015); and to Trechitae by Maddison et al. (2019).

Synonyms
Synonymy list
YearName and author
1827Bembidiina Stephens
1827Bembidiini Stephens
1861Bembidiini LeConte p. 35
1864Bembidiides Motschulsky
2004Bembidiinae Hayashi et al. p. 64
2011Bembidiina Bouchard et al. p. 113
2011Bembidiini Bouchard et al. p. 113
2011Bembidiini Tamutis et al.
2015Bembidiina Ortuño and Arillo p. 578
2015Bembidiini Ortuño and Arillo p. 579
2019Bembidiini Maddison et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
RankNameAuthor
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderAdephaga
familyCarabidae()
subfamilyTrechinae()
—Trechitae
tribeBembidiiniStephens 1827
tribeBembidiiniStephens 1827

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Tr. Bembidiini Stephens 1827
show all | hide all
G. Asaphidion Des Gozis 1886
+
Asaphidion alaskanum Wickham 1919
Asaphidion yukonense Wickham 1919
G. Bembidion Latreille 1802
+
Subg. Bembidion (Bembidion) Latreille 1802
+
Bembidion (Bembidion) mutatum Gemminger and Harold 1868
Subg. Bembidion (Bracteon) Bedel 1879
+
Invalid names: Bembidion (Chrysobracteon) Netolitzky 1914 [synonym]
Subg. Bembidion (Diplocampa) Bedel 1896
+
Subg. †Bembidion (Eodontium) Schmidt and Michalik 2017
+
†Bembidion (Eodontium) bukejsi Schmidt and Michalik 2017
Subg. Bembidion (Eupetedromus) Netolitzky 1911
+
†Bembidion (Eupetedromus) alekseevi Schmidt and Michalik 2017
Subg. Bembidion (Furcacampa) Netolitzky 1931
+
Subg. Bembidion (Hydrium) LeConte 1847
+
Invalid names: Bembidion (Eurytrachelus) Motschulsky 1850 [synonym]
Subg. Bembidion (Melomalus) Casey 1918
+
Subg. Bembidion (Ochthedromus) LeConte 1847
+
Subg. Bembidion (Philochthemphanes) Netolitzky 1943
+
†Bembidion (Philochthemphanes) christelae Ortuño and Arillo 2010
Invalid names: Bembidion (Archaeophilochthus) Ortuño and Arillo 2010 [synonym]
Subg. Bembidion (Phyla) Motschulsky 1844
+
Bembidion (Phyla) obtusum Audinet-Serville 1821
Subg. Bembidion (Plataphus) Motschulsky 1864
+
Bembidion (Plataphus) breve Motschulsky 1845
Invalid names: Bembidion (Plataphodes) Ganglbauer 1891 [synonym]
Subg. Bembidion (Princidium) Motschulsky 1864
+
Subg. Bembidion (Semicampa) Netolitzky 1910
+
Subg. Bembidion (Terminophanes) Müller-Motzfeld 1998
+
Subg. Bembidion (Testedium) Motschulsky 1864
+
Bembidion aeneum Germar 1824
†Bembidion damnosum Scudder 1900
†Bembidion everestae Pierce 1944
†Bembidion festivum Zhang et al. 1994
Bembidion gilvipes Sturm 1825
Bembidion lunatum Duftschmid 1812
Bembidion minimum Fabricius 1792
Bembidion obliquum Sturm 1825
†Bembidion strenum Zhang 1989
Bembidion virens Gyllenhal 1827
G. Ocys Stephens 1828
+
Ocys harpaloides Audinet-Serville 1821
G. Sinechostictus Motschulsky 1864
+
Invalid names: Bembidiina Stephens 1827 [invalid subgroup], Tachyina Motschulsky 1862 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Priabonian to the top of the Holocene or 37.71000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 33.9 Ma

Collections (94 total)


Time interval Ma Country or state Original ID and collection number
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Bembidion alekseevi (139928) Bembidion bukejsi (152022) Bembidion christelae (123462)
Burdigalian20.45 - 15.98USA (California) Bembidion davidae (161515) Bembidion everestae (161516)
Burdigalian20.45 - 15.98China (Shandong) Bembidion festivum (138946) Bembidion strenum (128552)
Late/Upper Miocene11.63 - 5.333Russian Federation Bembidion sp. 1 (153434) Bembidion sp. 4, Bembidion sp. 5, Bembidion sp. 1, Bembidion sp. 2, Bembidion sp. 3, Bembidion sp. (113713)
Messinian7.246 - 5.333USA (Alaska) Bembidion sp., Bembidion sp., Bembidion spp. (123352)
Piacenzian3.6 - 2.58USA (Alaska) Asaphidion yukonense (162793)
Gelasian2.58 - 1.8Greenland Bembidion mckinleyi (160159) Bembidion spp., Bembidion dyschirinum, Bembidion sordidum (160108) Bembidion spp., Bembidion fellmanni (160152) Bembidion spp., Bembidion fellmanni, Bembidion mckinleyi, Bembidion sordidum (160151) Bembidion spp., Bembidion levettei, Bembidion velox, Bembidion lampros, Bembidion yukonum (160153) Bembidion spp., Bembidion sordidum (160147) Bembidion spp., Bembidion sp., Asaphidion alaskanum, Bembidion spp., Bembidion levettei, Bembidion lapponicum, Bembidion balli, Bembidion vitiosum, Bembidion dyschirinum, Bembidion fellmanni, Bembidion difficile, Bembidion arcticum, Bembidion planatum, Bembidion planiusculum, Bembidion gebleri, Bembidion salebratum, Bembidion mckinleyi, Bembidion grapii, Bembidion yukonum, Bembidion bimaculatum, Bembidion sordidum (160109) Bembidion spp., Bembidion sp., Bembidion spp., Bembidion velox, Bembidion dyschirinum, Bembidion fellmanni, Bembidion arcticum, Bembidion planiusculum, Bembidion gebleri, Bembidion lenae, Bembidion yukonum, Bembidion sordidum (160146) Bembidion spp., Bembidion spp., Bembidion levettei, Bembidion alaskense, Bembidion dyschirinum, Bembidion fellmanni, Bembidion planatum, Bembidion planiusculum, Bembidion gebleri, Bembidion mckinleyi, Bembidion yukonum, Bembidion bimaculatum, Bembidion sordidum (160144)
Early/Lower Pleistocene2.58 - 0.774Russian Federation Bembidion spp. (163069 163089)
Chibanian0.774 - 0.129Russian Federation Bembidion sp. (163091 163095)
Chibanian0.774 - 0.129Japan (Kumamoto) Bembidion sp. (164901)
Chibanian0.774 - 0.129Japan Bembidion sp. (165704 165705 165707 165709)
Middle Pleistocene0.774 - 0.129Japan (Kumamoto) Bembidiinae indet. (165716)
Middle Pleistocene0.774 - 0.129United Kingdom Bembidion doris, Bembidion articulatum, Bembidion octomaculatum (168266) Bembidion doris, Bembidion punctulatum, Bembidion tetracolum, Bembidion elongatum, Bembidion assimile (168264) Bembidion lampros, Bembidion doris, Bembidion elongatum, Bembidion assimile, Bembidion minimum, Bembidion articulatum, Bembidion obtusum (168265) Bembidion lampros, Bembidion doris, Bembidion punctulatum, Bembidion tetracolum, Bembidion elongatum, Bembidion quadrimaculatum (168267) Bembidion lampros, Bembidion doris, Bembidion varium, Bembidion semipunctatum, Bembidion tetracolum, Bembidion elongatum, Bembidion gilvipes, Bembidion biguttatum (168270) Bembidion lampros, Bembidion gilvipes, Bembidion assimile, Bembidion minimum, Bembidion articulatum, Bembidium harpaloides (168268) Bembidion lampros, Bembidion semipunctatum, Bembidion assimile, Bembidion articulatum (168269) Bembidion lampros, Bembidion varium, Bembidion elongatum, Bembidion assimile (168272) Bembidion lunatum, Bembidion elongatum, Bembidion gilvipes, Bembidion assimile (168271)
Middle Pleistocene0.774 - 0.129United Kingdom (Warwickshire) Bembidion assimile (169391 169392 169393) Bembidion sp., Bembidion assimile, Bembidion schueppeli (169388)
Late/Upper Pleistocene0.129 - 0.0117Japan Bembidiinae indet. (166082)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Bembidium tibiale, Bembidium bipunctatum (153476)
Late/Upper Pleistocene0.129 - 0.0117USA (Colorado) Bembidion bimaculatum, Bembidion coloradense, Bembidion constricticolle (169336) Bembidion sp. (169333) Bembidion sp., Bembidion breve (169347 169350)
Late/Upper Pleistocene0.129 - 0.0117Canada (Ontario) Bembidion balli (174230) Bembidion sp. 1, Bembidion sp. 2, Bembidion balli (174229) Bembidium damnosum (152577)
Late/Upper Pleistocene0.129 - 0.0117USA (Wisconsin) Bembidion sp., Bembidion mutatum, Asaphidion yukonense, Bembidion grapei (169406)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation Bembidion sp. (153443 153444 153446 163101 163102 163104 163106 163110 173201 173202 173205 173206 173208 173216)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Worcestershire) Bembidion bipunctatum (163075) Bembidion spp., Bembidion lampros, Bembidion bipunctatum, Bembidion obliquum, Bembidion virens, Bembidion femoratum, Bembidion lunatum, Bembidion aeneum (163070)
Late/Upper Pleistocene0.129 - 0.0117USA (Washington) Bembidion fortestriatum (169379) Bembidion haruspex, Bembidion planiusculum (169374) Bembidion planiusculum (169365) Bembidion rusticum (169359) Bembidion sp. (169360 169370 169376 169378) Bembidion spp., Bembidion bimaculatum, Bembidion inaequale, Bembidion nigripes (157092 167662)
Late/Upper Pleistocene0.129 - 0.0117Canada (British Columbia) Bembidion spp., Bembidion dorsale (167673)
Late/Upper Pleistocene0.129 - 0.0117USA (Pennsylvania) Bembidion fortestriatum (137209 137212) Bembidion mutatum (137210) Bembidion versicolor (137211)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation (Astrakhan) Bembidion sp., Bembidion minimum (125794)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (England) Bembidion doris (164906 164909) Bembidion doris, Bembidion guttula (164905) Bembidion guttula (164904)
Tarantian0.129 - 0.0117USA (Pennsylvania) Bembidion anguliferum group (137201 137204) Bembidion anguliferum group, Bembidion versicolor, Bembidion pseudocautum, Bembidion frontale (137206) Bembidion incertum group, Bembidion mutatum (137205)
Holocene0.0117 - 0.0USA (Hawaii) Bembidion ignicola, Bembidion pacificum (224098)