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Panephemeroptera

Insecta

Taxonomy
Panephemeroptera was named by Crampton (1928). It is extant.

It was assigned to Hydropalaeoptera by Staniczek et al. (2011); and to Euhydropalaeoptera by Sroka et al. (2014).

Subtaxa
Ephemerida (syn. Reticulata), Ephemeronta, Ephemerontoida, Misthodotida, Syntonopteroidea
Sister taxa
Ephemerida (syn. Reticulata), Ephemeronta, Ephemerontoida

Synonyms
Synonymy list
YearName and author
1928Panephemeroptera Crampton
2007Ephemerontoida Willmann p. 112
2007Ephemeronta Willmann p. 113
2007Misthodotida Willmann p. 115
2011Panephemeroptera Staniczek et al.
2014Panephemeroptera Sroka et al. p. 979

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
RankNameAuthor
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
infraclassHydropalaeopteraRohdendorf 1969
—Euhydropalaeoptera
—PanephemeropteraCrampton 1928
—PanephemeropteraCrampton 1928

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Panephemeroptera Crampton 1928
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Unr. Ephemerida Latreille 1810 [mayfly]
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Unr. Heptabranchia Willmann 2007
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Or. †Coxoplectoptera Staniczek et al. 2011
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Fm. †Mickoleitiidae Staniczek et al. 2011
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G. †Mesogenesia Tshernova 1977
G. †Mickoleitia Staniczek et al. 2011
G. †Cretereisma Willmann 2007
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†Cretereisma antiqua Willmann 2007
†Cretereisma schwickertorum Willmann 2007
Or. Ephemeroptera Hyatt and Arms 1890 [mayfly]
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Fm. †Aenigmephemeridae Tshernova 1968
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G. †Aenigmephemera Tshernova 1968
Fm. †Babidae Kluge et al. 2006
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G. †Baba Kluge et al. 2006
G. †Clephemera Lin 1986
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†Clephemera clava Lin 1986
G. †Durlophlebia Sinitshenkova and Coram 2002
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†Durlophlebia radleyi Sinitshenkova and Coram 2002
Fm. †Epeoromimidae Tshernova 1969
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G. †Epeoromimus Tshernova 1969
G. †Foliomimus Sinitshenkova 1985
Unr. †Euplectoptera Tillyard 1932
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Unr. Anteritorna Kluge 1993
Fm. †Fuyoidae Zhang and Kluge 2007
Unr. Posteritorna Kluge et al. 1995
Invalid names: Aphelophlebodes Pierce 1945 [nomen dubium], Aphelophlebodes stocki Pierce 1945 [nomen dubium]
G. †Libanoephemera Azar et al. 2019
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G. †Parabaetis Haupt 1956
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†Parabaetis eocaenicus Haupt 1956
Subor. Schistonota McCafferty and Edmunds 1979
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Superfm. Ephemeroidea Latreille 1810
G. †Schistonotorum Jarzembowski and Wang 2019
Invalid names: Baetoidea Leach 1815 [empty], Leptophlebioidea Banks 1900 [empty]
Fm. †Triplosobidae Handlirsch 1906
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G. †Triplosoba Handlirsch 1906
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Invalid names: Blanchardia Brongniart 1894 [replaced]
G. †Turfanerella Demoulin 1954
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†Turfanerella tingi Ping 1935
Invalid names: Alulata Willmann 2007 [invalid subgroup], Aphelophlebia Pierce 1945 [synonym], Aphelophlebodidae Pierce 1945 [invalid subgroup], Costalimella nordestina Martins-Neto 1996 [nomen dubium], Ephemera oeningensis Heer 1864 [nomen nudum], Heptagenioidea [empty], Metretopus henningseni Demoulin 1965 [nomen dubium], Plectoptera Packard 1886 [synonym], Protephemeroidea Handlirsch 1906 [synonym], Triangulifera Willmann 2007 [synonym]
Fm. †Mesonetidae Tshernova 1969
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G. †Clavineta Sinitshenkova 1991
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†Clavineta brevinodia Huang et al. 2011
†Clavineta cantabilis Sinitshenkova 1991
†Clavineta citima Sinitshenkova 2000
†Clavineta excavata Huang et al. 2011
†Clavineta eximia Zhang 2006
†Clavineta transbaikalica Sinitshenkova 2000
G. †Furvoneta Sinitshenkova 1990
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†Furvoneta domefacta Sinitshenkova 1991
†Furvoneta khasurtensis Sinitshenkova 2017
†Furvoneta lata Sinitshenkova 1976
†Furvoneta lucida Sinitshenkova 2002
†Furvoneta relicta Zhang 2006
†Furvoneta sobria Sinitshenkova 2002
†Furvoneta undina Sinitshenkova 1976
G. †Mesoneta Brauer et al. 1889
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†Mesoneta antiqua Brauer et al. 1889
†Mesoneta beipiaoensis Wang 1980
†Mesoneta deusta Sinitshenkova 2000
†Mesoneta magna Sinitshenkova 1985
†Mesoneta minuta Sinitshenkova 2000
†Mesoneta mongolica Sinitshenkova 1989
†Mesoneta picta Sinitshenkova 2000
†Mesoneta triassica Sinitshenkova 2000
†Mesoneta tushilgae Sinitshenkova 1989
†Mesoneta uralensis Sharov 1948
†Mesoneta utriculata Sinitshenkova 1985
†Mesoneta zolensis Sinitshenkova 1990
Invalid names: Triassomachilis Sharov 1948 [synonym]
Invalid names: Triassomachilidae Sharov 1948 [synonym]
G. †Minorella Sinitshenkova and Marchal-Papier 2005
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†Minorella virgata Sinitshenkova and Marchal-Papier 2005
Subor. †Permoplectoptera Tillyard 1932
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Superfm. †Litophlebioidea Hubbard and Riek 1978
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Fm. †Litophlebiidae Hubbard and Riek 1978
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G. †Litophlebia Hubbard and Riek 1978
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Invalid names: Xenophlebia Riek 1976 [replaced]
G. †Triassolitophlebia Sinitshenkova and Aristov 2015
Invalid names: Xenophlebiidae Riek 1976 [replaced]
Fm. †Tintorinidae Krzemiński and Lombardo 2001
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G. †Tintorina Krzemiński and Lombardo 2001
Invalid names: Xenophlebioidea Riek 1976 [replaced]
Superfm. †Mesephemeroidea Lameere 1917
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Fm. †Mesephemeridae Lameere 1917
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G. †Mesephemera Handlirsch 1906
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Invalid names: Tineites Germar 1842 [replaced]
G. †Montralia Vía and Calzada 1987
G. †Palinephemera Lin 1986
Fm. †Palingeniopsidae Martynov 1938
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G. †Palingeniopsis Martynov 1932
Fm. †Sharephemeridae Sinitshenkova 2002
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G. †Hammephemera Sinitshenkova 2011
G. †Jurassephemera Zhang et al. 2022
G. †Sharephemera Sinitshenkova 2002
G. †Tunephemera Sinitshenkova 2013
Fm. †Misthodotidae Tillyard 1932
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G. †Misthodotes Sellards 1909
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†Misthodotes biguttatus Tillyard 1932
†Misthodotes delicatulus Tillyard 1936
†Misthodotes dubius Sinitshenkova 2013
†Misthodotes edmundsi Carpenter 1979
†Misthodotes obtusus Sellards 1907
†Misthodotes ovalis Tillyard 1932
†Misthodotes sharovi Tshernova 1965
†Misthodotes stapfi Kinzelbach and Lutz 1984
†Misthodotes tshernovae Sinitshenkova and Vassilenko 2012
†Misthodotes visherensis Novokshonov et al. 2002
†Misthodotes zalesskyi Tshernova 1965
Invalid names: Dromeus Sellards 1907 [replaced], Eudoter Tillyard 1936 [synonym]
G. †Thuringopteryx Kuhn 1937
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†Thuringopteryx gimmi Kuhn 1937
G. †Triassodotes Sinitshenkova and Marchal-Papier 2005
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†Triassodotes rasnitsyni Sinitshenkova 2021
†Triassodotes vogesiacus Sinitshenkova and Marchal-Papier 2005
Invalid names: Arnulfias Willmann 2007 [nomen nudum], Eudoteridae Demoulin 1954 [synonym], Eurekter Willmann 2007 [nomen nudum]
Fm. †Protereismatidae Handlirsch 1919
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G. †Alexandrinia Sinitshenkova and Vassilenko 2012
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†Alexandrinia directa Carpenter 1979
†Alexandrinia gigantea Sinitshenkova and Vassilenko 2012
†Alexandrinia ipsa Sinitshenkova 2013
†Alexandrinia vitta Sinitshenkova 2013
G. †Kukalova Demoulin 1970
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†Kukalova americana Demoulin 1970
G. †Ponalex Sinitshenkova and Aristov 2012
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†Ponalex maximus Sinitshenkova and Aristov 2012
G. †Protereisma Sellards 1907
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†Protereisma apicalis Martynov 1928
†Protereisma arcuatum Sellards 1907
†Protereisma elongatum Sellards 1907
†Protereisma insigne Tillyard 1932
†Protereisma latum Sellards 1907
†Protereisma permianum Sellards 1907
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Invalid names: Prodromus rectus Sellards 1907 [synonym], Protechma acuminatum Sellards 1907 [synonym], Protereisma gracile Sellards 1907 [synonym], Protereisma minus Sellards 1907 [synonym]
†Protereisma rossenrayensis Guthörl 1965
†Protereisma sellardsi Tillyard 1932
†Protereisma uralicum Zalessky 1946
Invalid names: Bantiska Sellards 1907 [synonym], Esca Sellards 1909 [synonym], Loxophlebia Martynov 1928 [synonym], Mecus Sellards 1909 [synonym], Pinctodia Sellards 1907 [synonym], Pinctodia curta Sellards 1907 [nomen dubium], Prodromus Sellards 1907 [synonym], Protechma Sellards 1907 [synonym], Recter Sellards 1907 [synonym], Rekter Sellards 1907 [synonym], Rekter extensus Sellards 1907 [nomen dubium], Scopus Sellards 1907 [synonym]
Invalid names: Kukalovidae Demoulin 1970 [synonym], Therates planus Sellards 1907 [nomen vanum]
Superfm. †Protereismatoidea DeMoulin 1958
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Fm. †Jarmilidae Demoulin 1970
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G. †Jarmila Demoulin 1970
Fm. †Oboriphlebiidae Hubbard and Kukalová-Peck 1980
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G. †Oboriphlebia Hubbard and Kukalová-Peck 1980
Invalid names: Cretereismatidae Willmann 2007 [empty], Mesephemeriformia Willmann 2007 [invalid subgroup]
G. †Phthartus Handlirsch 1904
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†Phthartus africanus Haughton 1924
†Phthartus netschajevi Handlirsch 1904
†Phthartus rossicus Handlirsch 1904
Unr. †Sinebranchia Sroka and Staniczek 2022
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Fm. †Mesoplectopteridae Demoulin 1955
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G. †Mesoplectopteron Handlirsch 1918
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†Mesoplectopteron longipes Handlirsch 1918
Fm. †Triassoephemeridae Sinitshenkova and Marchal-Papier 2005
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G. †Triassoephemera Sinitshenkova and Marchal-Papier 2005
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†Triassoephemera punctata Sinitshenkova and Marchal-Papier 2005
Fm. †Vogesonymphidae Sinitshenkova and Marchal-Papier 2005
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G. †Khungtukunia Sinitshenkova 2013
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†Khungtukunia sibirica Sinitshenkova 2013
G. †Palegonympha Sinitshenkova and Aristov 2015
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†Palegonympha triassica Sinitshenkova and Aristov 2015
G. †Vogesonympha Sinitshenkova and Marchal-Papier 2005
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†Vogesonympha ludovici Sinitshenkova and Marchal-Papier 2005
Fm. †Toxodotidae Sinitshenkova and Marchal-Papier 2005
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G. †Toxodotes Sinitshenkova and Marchal-Papier 2005
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†Toxodotes coloratus Sinitshenkova and Marchal-Papier 2005
Fm. †Triassomanthidae Sinitshenkova and Marchal-Papier 2005
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G. †Triassomanthus Sinitshenkova and Marchal-Papier 2005
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†Triassomanthus parvulus Sinitshenkova and Marchal-Papier 2005
Fm. †Voltziaephemeridae Sinitshenkova and Marchal-Papier 2005
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G. †Voltziaephemera Sinitshenkova and Marchal-Papier 2005
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†Voltziaephemera fossoria Sinitshenkova and Marchal-Papier 2005
G. †Warungata Sroka and Prokop 2024
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†Warungata peterjelli Sroka and Prokop 2024
Invalid names: Reticulata Willmann 2007 [synonym]
Superfm. †Syntonopteroidea Handlirsch 1911
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Fm. †Miracopteridae Novokshonov 1993
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G. †Miracopteron Novokshonov 1993
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†Miracopteron mirabile Novokshonov 1993
Fm. †Syntonopteridae Handlirsch 1911
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G. †Anglolithoneura Prokop et al. 2010
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†Anglolithoneura magnifica Prokop et al. 2010
G. †Anniedarwinia Beckemeyer and Engel 2011
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†Anniedarwinia alabamensis Beckemeyer and Engel 2011
G. †Carrizoneura Prokop and Kukalová-Peck 2017
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†Carrizoneura carpenteri Prokop and Kukalová-Peck 2017
G. †Gallolithoneura Garrouste et al. 2009
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†Gallolithoneura butchlii Garrouste et al. 2009
G. †Lithoneura Carpenter 1938
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†Lithoneura clayesi Kukalová-Peck 1985
†Lithoneura lameerei Carpenter 1938
†Lithoneura mirifica Carpenter 1943
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Invalid names: Lithoneura carpenteri Richardson 1956 [synonym]
†Lithoneura piecko Kukalová-Peck 1985
G. †Syntonoptera Handlirsch 1911
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†Syntonoptera schucherti Handlirsch 1911
Invalid names: Lithoneuridae Sroka et al. 2014 [empty]
Invalid names: Ephemeronta Willmann 2007 [invalid subgroup], Ephemerontoida Willmann 2007 [invalid subgroup], Misthodotida Willmann 2007 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Westphalian A to the top of the Piacenzian or 318.70000 to 2.58000 Ma
Minimum age of oldest fossil (stem group age): 316.9 Ma

Collections (355 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Westphalian A318.7 - 316.9United Kingdom (England) Anglolithoneura magnifica (102655)
Westphalian318.7 - 307.4USA (Alabama) Anniedarwinia alabamensis (81525)
Westphalian D309.8 - 307.4USA (Illinois) Lithoneura carpenteri (112638) Lithoneura clayesi (151853) Lithoneura lameerei (114453) Lithoneura mirifica (151849) Lithoneura piecko (102530) Syntonoptera schucherti (112630 122493) Syntonoptera schucherti, Lithoneura mirifica (133734)
Stephanian307.4 - 300.3France (Saône-et-Loire) Ephemeroptera indet. (236920)
Missourian306.0 - 303.7USA (New Mexico) Reticulata americana (215691)
Gzhelian303.7 - 298.9France Blanchardia pulchella (103635)
Virgilian303.7 - 298.9USA (Texas) Reticulata sp. (50197)
Asselian298.9 - 293.52Czechia Syntonopteroidea indet., Protereisma nymph 3, Protereisma nymph 4, Protereisma nymph 5, Protereisma nymph 2, Ephemeroptera nymph 6 (114186)
Asselian298.9 - 293.52Germany (Rheinland-Pfalz) Misthodotidae indet. (139349)
Asselian298.9 - 293.52USA (New Mexico) Carrizoneura carpenteri (122481)
Artinskian290.1 - 283.3USA (Oklahoma) Misthodotes ovalis, Protereisma directum, Misthodotes edmundsi, Protereisma nymph 1 (104824) Protereisma latum (104826) Protereisma sp., Protereisma arcuatum (107405)
Artinskian290.1 - 283.3USA (Kansas) Protereisma insigne (114152) Protereisma insigne, Misthodotes obtusus, Misthodotes ovalis (114151) Protereisma latum, Protereisma permianum, Protechma acuminatum, Prodromus rectus, Scopus gracilis, Protereisma minus, Bantiska elongata, Rekter arcuatus, Dromeus obtusus, Pinctodia curta, Rekter extensus, Therates planus (21979) Protereisma latum, Protereisma permianum, Protereisma arcuatum, Protereisma insigne (111319) Protereisma latum, Protereisma permianum, Protereisma elongatum, Protereisma arcuatum, Misthodotes obtusus (111320) Protereisma sp., Protereisma latum, Protereisma permianum, Protereisma minus, Protereisma gracile, Protereisma sellardsi, Protereisma elongatum, Protereisma arcuatum, Protereisma insigne, Misthodotes obtusus, Eudoter delicatulus, Misthodotes biguttatus, Misthodotes ovalis (111219) Protereismidae indet., Protereisma permianum (151412)
Kungurian283.3 - 274.4Russian Federation (Suksun) Misthodotes sharovi (124241) Misthodotes sp., Misthodotes zalesskyi, Misthodotes sharovi (108206)
Kungurian283.3 - 274.4France Syntonopteridae indet. (117603)
Kungurian283.3 - 274.4Russian Federation Miracopteron mirabile (107787 114207) Misthodotes sharovi (196228) Protereisma uralicum (161939)
Kungurian283.3 - 274.4Russian Federation (Perm) Misthodotes visherensis (117254)
Roadian274.4 - 266.9Russian Federation (Arkangelsk) Palingeniopsis praecox (111189) Protereismidae indet. (104993)
Roadian274.4 - 266.9Russian Federation (Kazan) Protereismidae indet. (116803)
Roadian274.4 - 266.9Russian Federation Loxophlebia apicalis (116456)
Capitanian264.28 - 259.51Russian Federation (Orenburg) Phthartus netschajevi, Phthartus rossicus (127931)