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Helophoridae
Taxonomy
Helophorida was named by Leach (1815). It is extant.
It was reranked as the tribe Helophorina by Sahlberg (1880); it was reranked as the subfamily Helophorinae by Lawrence and Newton (1995) and Bouchard et al. (2011); it was corrected as Helophoridae by Short and Fikáček (2013) and Cai et al. (2022).
It was assigned to Hydrophilidae by Sahlberg (1880), Lawrence and Newton (1995) and Bouchard et al. (2011); and to Hydrophiloidea by Short and Fikáček (2013) and Cai et al. (2022).
It was reranked as the tribe Helophorina by Sahlberg (1880); it was reranked as the subfamily Helophorinae by Lawrence and Newton (1995) and Bouchard et al. (2011); it was corrected as Helophoridae by Short and Fikáček (2013) and Cai et al. (2022).
It was assigned to Hydrophilidae by Sahlberg (1880), Lawrence and Newton (1995) and Bouchard et al. (2011); and to Hydrophiloidea by Short and Fikáček (2013) and Cai et al. (2022).
Subtaxa
Synonymy list
Year | Name and author |
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1880 | Helophorina Sahlberg p. 61 |
1995 | Helophorinae Lawrence and Newton, Jr. |
2011 | Helophorinae Bouchard et al. p. 154 |
2013 | Helophoridae Short and Fikáček |
2022 | Helophoridae Cai et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Helophoridae Leach 1815
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G. Helophorus Fabricius 1775
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Subg. Helophorus (Atracthelophorus) Kuwert 1890
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Helophorus (Atracthelophorus) aspericollis Angus 1973
Helophorus (Atracthelophorus) glacialis Villa and Villa 1833
Subg. Helophorus (Gephelophorus) Sharp 1915
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Helophorus (Gephelophorus) sibiricus Motschulsky 1860
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Invalid names: Helophorus (Gephelophorus) lomnicki d'Orchymont 1927 [synonym], Helophorus (Trichelophorus) pleistocenicus Lomnicki 1894 [synonym]
Subg. Helophorus (Meghelophorus) Kuwert 1886
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Helophorus (Meghelophorus) aquaticus Linnaeus 1758
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Invalid names: Helophorus (Praehelephorus) dzieduszyckii Lomnicki 1894 [synonym]
Helophorus (Meghelophorus) grandis Illiger 1798
Helophorus (Meghelophorus) khnzoriani Angus 1970
Invalid names: Helophorus (Praehelephorus) Lomnicki 1894 [synonym]
Subg. †Helophorus (Mesohelophorus) Ponomarenko 1977
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†Helophorus (Mesohelophorus) palaeosibiricus Fikáček et al. 2012
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Invalid names: Mesohelophorus sibiricus Ponomarenko 1977 [replaced]
Subg. †Helophorus (Mesosperchus) Ponomarenko 1977
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†Helophorus (Mesosperchus) costalis Ponomarenko 2014
†Helophorus (Mesosperchus) gracilis Prokin et al. 2010
†Helophorus (Mesosperchus) inceptivus Fikáček et al. 2012
†Helophorus (Mesosperchus) tarsalis Ponomarenko 1977
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Invalid names: Mesohelophorus elongatus Ponomarenko 1990 [synonym]
†Helophorus (Mesosperchus) yixianus Fikáček et al. 2012
Subg. Helophorus (Orphelophorus) d'Orchymont 1927
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Helophorus (Orphelophorus) obscurellus Poppius 1907
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Invalid names: Helophorus (Orphelophorus) wandereri d'Orchymont 1927 [synonym]
Subg. Helophorus (Rhopalhelophorus) Kuwert 1886
Subg. Helophorus (Trichelophorus) Kuwert 1886
Helophorus auricollis Eschscholtz 1822
Helophorus brevipalpis Bedel 1881
Helophorus browni McCorkle 1973
Helophorus discrepans Rey 1885
Helophorus eclectus d'Orchymont 1945
Helophorus lacustris Le Conte 1850
Helophorus nubilus Fabricius 1777
Helophorus oblongus Le Conte 1850
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Invalid names: Helophorus (Rhopalhelophorus) kuwerti Lomnicki 1894 [synonym]
Helophorus orientalis Motschulsky 1860
Helophorus sempervarians Angus 1970
Helophorus splendidus Sahlberg 1880
Helophorus tuberculatus Gyllenhal 1808
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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Source: c = class, p = phylum | |||||
References: Kiessling 2004, Bush and Bambach 2015 |
Age range
Maximum range based only on fossils: base of the Tithonian to the top of the Late/Upper Pleistocene or 152.10000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 145.0 Ma
Minimum age of oldest fossil (stem group age): 145.0 Ma
Collections (74 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Tithonian | Russian Federation | Mesosperchus tarsalis (125408 138150) | |
Tithonian | Mongolia | Helophorus inceptivus (131803) Helophorus spp., Helophorus costalis (104748) Helophorus spp., Helophorus inceptivus, Helophorus costalis (121394) | |
Tithonian | Russian Federation (Chita) | Mesohelophorus elongatus (125407) Mesosperchus tarsalis (119424 123197 152174) Mesosperchus tarsalis, Mesohelophorus elongatus (114307) | |
Hauterivian | Russian Federation (Chita) | Mesosperchus tarsalis (151559) | |
Late/Upper Barremian | China (Liaoning) | Hydrophilopsia gracilis, Helophorus yixianus (111716) | |
Barremian | Russian Federation (Chita) | Mesosperchus tarsalis (151642) | |
Aptian | Russian Federation (Buryatia) | Mesohelophorus sibiricus (105536 105969) | |
Early/Lower Aptian | Russian Federation (Chita) | Mesosperchus tarsalis (152173) | |
Aquitanian - Burdigalian | Russian Federation | Helophorus sibiricus (117611) | |
Late/Upper Miocene | Russian Federation | Helophorus tuberculatus (153434) | |
Messinian | USA (Alaska) | Helophorus sp. (123352) | |
Gelasian | Greenland | Helophorus khnzoriani (160143) Helophorus spp., Helophorus khnzoriani (160144) Helophorus spp., Helophorus tuberculatus, Helophorus khnzoriani (160109) | |
Early/Lower Pleistocene | Russian Federation | Helophorus splendidus (163069) | |
Chibanian | Russian Federation | Helophorus obscurellus (163093) Helophorus splendidus (163094 163097) | |
Middle Pleistocene | United Kingdom (Warwickshire) | Helophorus sp. (169389) | |
Middle Pleistocene | United Kingdom | Helophorus sp. (168265 168266 168267 168270 168271) Helophorus spp. (168264 168269) | |
Late/Upper Pleistocene | Russian Federation | Helophorus splendidus (153437 153439 153443 153446 173208) | |
Late/Upper Pleistocene | Ukraine | Helophorus pleistocenicus, Helophorus dzieduszyckii, Helophorus praenanus, Helophorus polonicus, Helophorus kuwerti, Helophorus aquaticus, Helophorus lomnicki (153476) Helophorus sp., Helophorus sibiricus, Helophorus aquaticus, Helophorus praenanus, Helophorus grandis, Helophorus glacialis, Helophorus aspericollis, Helophorus orientalis, Helophorus browni, Helophorus discrepans (113753) | |
Late/Upper Pleistocene | USA (Washington) | Helophorus lacustris (169359 169363) Helophorus oregonensis (169370 169372 169374 169376) Helophorus oregonensis, Helophorus lacustris (169366) Helophorus oregonensis, Helophorus sp., Helophorus lacustris (169361) Helophorus sp. (169362) Helophorus sp., Helophorus lacustris (169360) Helophorus spp. (157092 167662 169375 169379) | |
Late/Upper Pleistocene | USA (Colorado) | Helophorus auricollis (169350) Helophorus eclectus (169347) | |
Late/Upper Pleistocene | Germany | Helophorus lomnicki, Helophorus wandereri (156167) | |
Late/Upper Pleistocene | United Kingdom (England) | Helophorus brevipalpis (164903 164920) | |
Late/Upper Pleistocene | United Kingdom (Worcestershire) | Helophorus aquaticus, Helophorus nubilus, Helophorus brevipalpis (163070) | |
Late/Upper Pleistocene | USA (Alaska) | Helophorus splendidus (169385) | |
Late/Upper Pleistocene | USA (Pennsylvania) | Helophorus sempervarians (137207 137210 137211 137213) Helophorus sp., Helophorus sempervarians (137208) Helophorus tuberculatus (137209) Helophorus tuberculatus, Helophorus sempervarians (137212) | |
Late/Upper Pleistocene | Canada (British Columbia) | Helophorus sp. (167675) | |
Tarantian | USA (Pennsylvania) | Helophorus sp. (137201 137204) Helophorus sp., Helophorus sempervarians (137205 137206) |