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Aphodiinae

Insecta - Coleoptera - Scarabaeidae

Synonymy list
YearName and author
1995Aphodiinae Lawrence and Newton, Jr.
2000Aphodiinae Krell p. 881
2003Aphodiinae Howden
2003Aphodiinae Wappler
2007Aphodiinae Nikolajev p. 122
2011Aphodiinae Bouchard et al. p. 239
2014Aphodiinae Nikolajev et al. p. 220
2016Aphodiinae Webster
2017Aphodiinae Tamutis et al.
2018Aphodiinae Bukejs and Alekseev
2021Aphodiinae Skelley
2022Aphodiinae Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
RankNameAuthor
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderPolyphagaEmery 1886
infraorderStaphyliniformia
superfamilyScarabaeoidea
familyScarabaeidae()
subfamilyAphodiinae()
subfamilyAphodiinae()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Aphodiinae Leach 1815
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Tr. Aphodiini Leach 1815
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G. Agoliinus Schmidt 1913
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Subtr. Aphodiina Leach 1815
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G. Aphodius Hellwig 1798
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Subg. Aphodius (Acrossus) Mulsant 1842
Subg. Aphodius (Agrilinus) Mulsant and Rey 1870
Subg. Aphodius (Nialus) Mulsant and Rey 1870
Subg. Aphodius (Plagiogonus) Mulsant 1842
†Aphodius aboriginalis Wickham 1912
†Aphodius anteactus Krell 2000
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Invalid names: Aphodius antiquus Heer 1847 [replaced]
†Aphodius boryslavicus Lomnicki 1894
†Aphodius bosniaskii Handlirsch 1907
†Aphodius brevipennis Heer 1862
Aphodius carpetanus Graells 1847
†Aphodius charauxi Piton 1940
Aphodius conspurcatus Linnaeus 1758
†Aphodius cretaceous Nikolajev 2008
Aphodius distinctus Müller 1776
Aphodius fimitarius Linnaeus 1758
†Aphodius florissantensis Wickham 1911
Aphodius fossor Linnaeus 1758
†Aphodius granarioides Wickham 1913
Aphodius granarius Linnaeus 1767
†Aphodius helvolus Statz 1952
Aphodius holdereri Reitter 1900
†Aphodius inundatus Wickham 1914
†Aphodius krantzi von Heyden and von Heyden 1866
†Aphodius laminicola Wickham 1910
†Aphodius mediaevus Wickham 1914
†Aphodius meyeri Heer 1847
Aphodius plagiatus Linnaeus 1767
†Aphodius praeemptor Wickham 1913
†Aphodius precursor Horn 1876
†Aphodius rhinocerontis Lomnicki 1894
Aphodius rufipes Linnaeus 1758 [night-flying dung beetle]
†Aphodius ruthenus Lomnicki 1894
†Aphodius schlickumi Statz 1952
†Aphodius senex Wickham 1914
†Aphodius shoshonis Wickham 1912
†Aphodius subater Lomnicki 1894
†Aphodius theobaldi Krell 2000
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Invalid names: Aphodius incertus Theobald 1937 [replaced]
†Aphodius vectis Krell 2019
G. Diapterna Horn 1891
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G. Heptaulacus Mulsant 1842
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Heptaulacus sus Herbst 1783
G. Caligodorus Gordon and Skelley 2007
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Caligodorus opacus Le Conte 1872
G. Dialytes Harold 1869
G. Dialytodius Gordon and Skelley 2007
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G. Pardalosus Gordon and Skelley 2007
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Pardalosus pardalis Le Conte 1857
G. Planolinoides Dellacasa and Dellacasa 2005
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Planolinoides duplex LeConte 1878
Tr. Eupariini Schmidt 1910
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G. Airapus Stebnicka and Howden 1996
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†Airapus lithuanicus Tamutis et al. 2017
G. Ataenius Harold 1867
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†Ataenius damzeni Bukejs and Alekseev 2018
†Ataenius europaeus Quiel 1911
†Ataenius patescens Scudder 1900
†Ataenius restructus Wickham 1912
G. Oxyomus Kirby and Spence 1828
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†Oxyomus nearcticus Wickham 1914
G. Saprosites Redtenbacher 1858
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†Saprosites cascus Britton 1960
†Saprosites succini Zang 1905
Tr. †Psammaegialiini Nikolajev et al. 2014
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G. †Psammaegialia Nikolajev et al. 2014
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†Psammaegialia abdita Nikolajev et al. 2014
†Psammaegialia zebrina Nikolajev et al. 2014
Tr. Psammodiini Mulsant 1842
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Subtr. Psammodiina Mulsant 1842
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G. Pleurophorus Mulsant 1842
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Pleurophorus caesus Creutzer 1796
G. Psammodius Fallen 1807
Invalid names: Pleurophorina Mulsant 1842 [synonym]
Subtr. Rhyssemina Pittinno and Marianni 1986
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G. Rhyssemus Mulsant 1842
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Rhyssemus germanus Linnaeus 1767
Tr. Rhyparini Schmidt 1910
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G. Leptorhyparus Howden 2003
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G. †Priscyparus Skelley 2021
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†Priscyparus quisqueyensis Skelley 2021
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Late/Upper Barremian to the top of the Late/Upper Pleistocene or 125.77000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 121.4 Ma

Collections (116 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Barremian125.77 - 121.4China (Nei Mongol) Psammaegialia abdita, Psammaegialia zebrina (103654)
Aptian121.4 - 113.2Russian Federation (Buryatia) Aphodius cretaceous (118216)
Selandian61.66 - 59.24France (Auvergne) Aphodius charauxi (113685)
Ypresian56.0 - 48.07United Kingdom (England) Saprosites cascus (126591)
MP 1348.07 - 41.03Germany (Eifel) Aphodiinae indet. 1, Aphodiinae indet. 2, Aphodiinae indet. 3 (138912) Aphodiinae indet. 2 (135644)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Airapus lithuanicus (194874) Aphodius succini (123221) Ataenius europaeus (123927)
Priabonian37.71 - 33.9Poland Airapus lithuanicus (124519) Ataenius damzeni (110737)
Priabonian37.71 - 33.9France Aphodius incertus (113709)
Priabonian37.71 - 33.9United Kingdom (England) Aphodius vectis (21976)
Chadronian37.0 - 33.9USA (Colorado) Aphodius aboriginalis, Aphodius shoshonis (106382) Aphodius florissantensis (106379) Aphodius laminicola (122975) Aphodius restructus (114150) Aphodius sp., Ataenius patescens, Aphodius aboriginalis, Aphodius shoshonis, Aphodius laminicola, Oxyomus nearcticus, Ataenius restructus, Aphodius mediaevus, Aphodius praeemptor, Aphodius senex (106384) Ataenius patescens, Aphodius aboriginalis, Aphodius laminicola, Aphodius praeemptor, Aphodius inundatus, Aphodius granarioides (113749) Ataenius patescens, Aphodius granarioides (124624) Ataenius patescens, Aphodius laminicola (106381)
Chattian27.3 - 23.04Germany (North Rhine-Westphalia) Aphodius schlickumi, Aphodius helvolus (122010)
Chattian27.3 - 23.04France Aphodius sp. (160861)
MP 3027.3 - 23.04Germany Aphodius krantzi (123945)
Early/Lower Miocene23.04 - 15.98Ukraine Aphodiinae indet. (133082)
Early/Lower Miocene23.04 - 15.98Japan Aphodius sp. (114003)
Burdigalian - Langhian20.45 - 13.82Dominican Republic Leptorhyparus quadricornis (146610) Priscyparus quisqueyensis, Leptorhyparus quadricornis (123857)
Sarmatian12.8 - 11.63Germany (Baden-Württemberg) Aphodius antiquus (115304) Aphodius brevipennis (117252) Aphodius meyeri (115305)
Messinian7.246 - 5.333Italy (Tuscany) Aphodius bosniaskii (113751)
Piacenzian3.6 - 2.58Germany (Niedersachsen) Aphodius rufipes (109196)
Gelasian2.58 - 1.8Greenland Aphodius sp. (160109)
Early/Lower Pleistocene2.58 - 0.774Japan Aphodius sp. (153617)
Early/Lower Pleistocene2.58 - 0.774Russian Federation Aphodius sp. (163069)
Irvingtonian1.4 - 0.21USA (Pennsylvania) Aphodius precursor (20479)
Chibanian0.774 - 0.129Russian Federation Aphodius sp. (163090 163091 163092 163093 163095 163096 163097 163098)
Chibanian0.774 - 0.129Japan Aphodius sp. (165700 165704)
Chibanian0.774 - 0.129Japan (Kumamoto) Aphodius sp. (164901)
Middle Pleistocene0.774 - 0.129United Kingdom Aphodius sp., Heptaulacus sus, Rhyssemus germanus, Pleurophorus caesus (168264) Aphodius spp. (168268 168270) Aphodius spp., Aphodius carpetanus, Aphodius conspurcatus, Heptaulacus sus (168265) Aphodius spp., Heptaulacus testudinarius (168271 168272) Aphodius spp., Heptaulacus testudinarius, Heptaulacus sus, Pleurophorus caesus (168267) Aphodius spp., Heptaulacus testudinarius, Rhyssemus germanus (168266) Aphodius spp., Pleurophorus caesus (168269)
Middle Pleistocene0.774 - 0.129United Kingdom (Warwickshire) Aphodius sp. (169388 169389 169393 169394 169395 169396 169399 169400) Aphodius sp., Aphodius sp. A (169391 169392)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Aphodius granarius, Aphodius subater, Aphodius rufipes, Aphodius rhinocerontis, Aphodius ruthenus, Aphodius boryslavicus (153476)
Late/Upper Pleistocene0.129 - 0.0117USA (Pennsylvania) Aphodius sp. (137208)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Worcestershire) Aphodius sp., Aphodius spp., Aphodius rufipes, Aphodius fossor, Aphodius distinctus, Aphodius plagiatus, Aphodius fimitarius, Aphodius sp. A (163070)
Late/Upper Pleistocene0.129 - 0.0117Japan Aphodius sp. (166083 166084)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation (Astrakhan) Aphodius sp., Aphodius sp., Aphodius sp., Aphodius sp., Aphodius sp. (125794)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (England) Aphodius sp. (164903 164919 164920)
Late/Upper Pleistocene0.129 - 0.0117USA (Colorado) Aphodius plutonicus, Dialytes sp. (169336) Aphodius sp., Aphodius duplex, Aphodius decipiens (169347) Aphodius spp. (169350) Aphodius spp., Ataenius sp., Aphodius decipiens (169353)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Oxfordshire) Aphodius holdereri (163112)
Late/Upper Pleistocene0.129 - 0.0117USA (Washington) Aphodius opacus (169360) Aphodius pectoralis (169361 169362) Aphodius sp. (169365 169366 169367 169371 169374) Aphodius sp., Aphodius pectoralis (169370) Aphodius spp. (169359) Diapterna hamata, Aphodiinae indet. (157092) Diapterna hamata, Pardalosus pardalis, Aphodiinae indet. (167662)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Northamptonshire) Aphodius holdereri (163111)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation Aphodius sp. (153436 153438 153439 153440 153441 153442 153443 153444 153445 153446 163104 163105 173201 173202 173203 173204 173208 173210)
Tarantian0.129 - 0.0117USA (Pennsylvania) Aphodius sp. (137201 137204 137205 137206)