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Titanochelon bacharidisi
Discussion
Named after Mr. Nikolaos Bacharidis, who discovered and excavated the type specimen
Taxonomy
Cheirogaster bacharidisi was named by Vlachos et al. (2014). Its type specimen is LGPUT EPN 100-199, a partial skeleton (an almost complete skeleton, missing most of the carapace), and it is a 3D body fossil. Its type locality is Epanomi (=EPN), which is in a Pliocene/Pliocene fluvial claystone in the Gonia Formation of Greece.
It was recombined as Titanochelon bacharidisi by Pérez-García and Vlachos (2014), Lujan et al. (2017), Vlachos (2021).
It was recombined as Titanochelon bacharidisi by Pérez-García and Vlachos (2014), Lujan et al. (2017), Vlachos (2021).
Synonymy list
| Year | Name and author |
|---|---|
| 2014 | Titanochelon bacharidisi Pérez-García and Vlachos |
| 2014 | Cheirogaster bacharidisi Vlachos et al. |
| 2017 | Titanochelon bacharidisi Lujan et al. |
| 2021 | Titanochelon bacharidisi Vlachos |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Titanochelon bacharidisi Vlachos et al. 2014
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Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| E. Vlachos 2021 | This species is a member of the derived geocheloninan clade Titanochelon, sharing several characters with other titanochelons, including presence of frontals narrower than prefrontals, a long crista supraoccipitalis, the presence of an elliptical to circular premaxillary pit, two suprapygals, the first surrounding the lenticular second one, absence of cervical scute, a convex to flat dorsal lip of the epiplastra, a humero-pectoral sulcus that is perpendicular to the axial plane in the medial region, but with a well-developed lateral change of curvature, very short pectoral scutes in the sagittal plane, but markedly expanded, toward the anterior region, laterally. Among other titanochelons, Titanochelon bacharidisi is diagnosed because of the position of the fossa orbitalis in the midway between the apertura narium externa and the posterior tip of the squamosal; a zygomatic arch that is thick and short; squamosal processes converging posteriorly toward the midline; extended ventral emargination, short gulars contacting the anterior part of the entoplastron, which is covered medially only by humeral scutes; and its posterior border coinciding with the humero-pectoral sulcus. This species was also characterized by the presence of at least one conical, bony, thigh tubercle on each side of the tail (found in anatomical position in the Nea Kallikratia material shown in Fig. 5d and isolated in the Epanomi holotype), similar to the morphology of the extant African spurred tortoise (Centrochelys sulcata). |
Measurements
No measurements are available
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| Source: f = family, c = class, subp = subphylum | |||||
| References: Carroll 1988, Ernst and Barbour 1989, Hendy et al. 2009 | |||||
Age range: base of the Ruscinian to the top of the Early/Lower Pliocene or 5.09000 to 3.60000 Ma
Collections (5 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Early/Lower Pliocene | Bulgaria (Yambol) | Testudo sp. (184410) | |
| Early/Lower Pliocene - Late/Upper Pliocene | Greece (Thessaloniki) | Cheirogaster bacharidisi (type locality: 182672 182673 182674) | |
| Ruscinian | Greece | Testudo sp. (36579) |