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Testudo marmorum
Discussion
MNHNPIK 3683a (lectotype by designation of Vlachos et al. 2019), a partial shell This species is named in Gaudry, A. 1862a. Résultats des fouilles exécutées en Grèce sous les auspices de l’Académie. Comptes Rendus de l’Académie des Sciences de Paris 52:297–300, which has priority
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Taxonomy
Testudo marmorum was named by Gaudry (1862). Its type specimen is MNHN PIK 3683a, a partial shell, and it is a 3D body fossil. Its type locality is Pikermi (old collection), which is in a Miocene fluvial horizon in Greece.
Sister species lacking formal opinion data
Synonymy list
| Year | Name and author |
|---|---|
| 1862 | Testudo marmorum Gaudry p. 635 figs. pl. 16 |
| 2016 | Testudo marmorum Lujan et al. |
| 2018 | Testudo marmorum Vlachos and Rabi p. 663 |
| 2019 | Testudo marmorum Vlachos et al. |
| 2021 | Testudo marmorum Vlachos |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Testudo marmorum Gaudry 1862
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Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| E. Vlachos et al. 2019 | Testudinidae based on the presence of a well-developed epiplastral lip, well-developed anterior and posterior buttresses, lateral coincidence between the costoperipheral sutures and the pleuromarginal sulci, presence of a supracaudal, and short pectoral scutes. Testudona based on the absence of contact between marginal 6 and pleural 3. Testudo based on the presence of a hinge between the hypoplastra and the xiphiplastra, at least in male adults. Differs from other species of Testudo by the following combination of characters: cervical scute short and narrow, vertebral 1 lyre-shaped, pleural 1 covers lateral parts of the nuchal, there is no coincidence between the anterior pleuromarginal sulci and costoperipheral sutures and the first two marginals expand on costal 1, suprapygal 1 is boomerang-shaped, embracing the smaller suprapygal 2, and gular scutes are short and cover the anterior part of the entoplastron. | |
| E. Vlachos 2021 | Testudo marmorum is clearly distinguished by other Testudo based on the following combination of characters: cervical scute is short and narrow, vertebral 1 is lyre-shaped, pleural 1 covers the lateral parts of the nuchal, there is no coincidence between the anterior pleuro-marginal sulci and the costo-peripheral suture and the first two marginals expand on the costal 1, suprapygal 1 is boomerang-shaped embracing the smaller suprapygal 2, and gular scutes are short and cover the anterior part of the entoplastron |
Measurements
No measurements are available
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| Source: f = family, c = class, subp = subphylum | |||||
| References: Hendy et al. 2009, Carroll 1988, Ernst and Barbour 1989 | |||||
Collections (11 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| MN 5 - MN 6 | Türkiye (Ankara) | Testudo graeca (68561) | |
| Sarmatian | Türkiye | Testudo sp. (56563) | |
| Late/Upper Miocene | Greece (Attica) | Testudo marmorum (type locality: 182754) Testudo sp. (202630) | |
| MN 11 | Greece | Testudo marmorum (73869) | |
| Turolian | Greece (Samos Island) | Testudo marmorum (205351) | |
| Turolian | Bulgaria | Testudo marmorum (208566 208567 208568 208569) | |
| Turolian | Greece (Euboea) | Testudo marmorum (202213) |