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Emys (pond turtle)
Taxonomy
Emys was named by Dumeril (1806). It is extant. It is the type genus of Emydidae, Emydinae.
It was assigned to Chelonia by Leidy (1865); to Chelonia by Leidy (1860) and Seeley (1881); to Paludinosa by Quenstedt (1882); to Testudinidae by Lydekker (1889); to Emydinae by Bickham (1975), Holman (2002) and van Dijk et al. (2014); and to Emydidae by Owen (1842), Leidy (1873), Cope (1875), Cope (1884), Hay (1902), Carroll (1988), Capasso Barbato and Gliozzi (1995), Frank and Ramus (1996), Vlachos and Delfino (2016), Vlachos (2018) and Vlachos (2021).
It was assigned to Chelonia by Leidy (1865); to Chelonia by Leidy (1860) and Seeley (1881); to Paludinosa by Quenstedt (1882); to Testudinidae by Lydekker (1889); to Emydinae by Bickham (1975), Holman (2002) and van Dijk et al. (2014); and to Emydidae by Owen (1842), Leidy (1873), Cope (1875), Cope (1884), Hay (1902), Carroll (1988), Capasso Barbato and Gliozzi (1995), Frank and Ramus (1996), Vlachos and Delfino (2016), Vlachos (2018) and Vlachos (2021).
Synonyms
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Synonymy list
Year | Name and author |
---|---|
1806 | Emys Dumeril |
1842 | Emys Owen p. 160 |
1860 | Emys Leidy p. 153 |
1860 | Emys namadicus Theobald p. 295 |
1865 | Emys Leidy p. 106 |
1873 | Emys Leidy p. 340 |
1875 | Emys Cope p. 53 |
1881 | Emys Seeley p. 620 |
1882 | Emys Quenstedt p. 149 |
1884 | Emys Cope |
1889 | Emys Lydekker p. 102 |
1902 | Emys Hay p. 447 |
1975 | Emys Bickham p. 203 |
1988 | Emys Carroll |
1995 | Emys Capasso Barbato and Gliozzi p. 238 |
1996 | Emys Frank and Ramus |
2002 | Emys Holman p. 440 |
2014 | Emys van Dijk et al. |
2016 | Emys Vlachos and Delfino |
2018 | Emys Vlachos |
2021 | Emys Vlachos |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. Emys Dumeril 1806 [pond turtle]
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†Emys cuvieri Gray 1831
Emys dhongoka Gray 1834 [Three-striped roofed turtle]
†Emys hordwellensis Seeley 1876
Emys kachuga Gray 1831 [Red-crowned roofed turtle]
†Emys lucii Gray 1831
Emys marmorata Baird and Girard 1852 [western pond turtle]
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Invalid names: Clemmys owyheensis Brattstrom and Sturn 1959 [synonym]
Emys orbicularis Linnaeus 1758 [European pond turtle]
Emys sigriz Michahelles 1829
Emys trinacris Fritz et al. 2005 [Sicilian pond turtle]
Emys vulgaris Gray 1831
†Emys wermuthi Mlynarski 1956
†Emys wyomingensis Leidy 1870
†Ocadia sinensis changwui Tao 1988
Invalid names: Emys namadicus Theobald 1860 [nomen nudum]
Diagnosis
Reference | Diagnosis | |
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E. Vlachos 2021 | Fossil emydids known from shell remains can be readily distinguished based on the presence of a wide costovertebral tunnel and wide rib heads, the absence of musk ducts and their foramina, and the expansion of the vertebral V onto the pygal bone. |
Measurements
No measurements are available
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Source: f = family, c = class, subp = subphylum | |||||
References: Carroll 1988, Hendy et al. 2009, Uetz 2005 |
Age range
Maximum range based only on fossils: base of the Puercan to the top of the Holocene or 66.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 60.9 Ma
Minimum age of oldest fossil (stem group age): 60.9 Ma
Collections (116 total)
Time interval | Ma | Country or state | Original ID and collection number |
---|---|---|---|
Puercan - Torrejonian | USA (New Mexico) | Emys sp. (131857) | |
Wasatchian | USA (Wyoming) | Emys sp. (180111) | |
Early/Lower Eocene | China (Jiangxi) | Emys sp. (182565) | |
Bridgerian | USA (Wyoming) | Chrysemys wyomingensis (16284) Emys wyomingensis (16211 128447 130809) | |
Middle Eocene | USA (Wyoming) | Emys wyomingensis (81455) | |
Bartonian | United Kingdom (England) | Chelonia indet. (39014) | |
Priabonian | United Kingdom (England) | Emys sp. (39643 39665 39667) | |
Late/Upper Eocene | United Kingdom (England) | Emys sp. (39662 40030) | |
MP 17A - MP 17B | United Kingdom (England) | Emys sp., Emys hordwellensis (28747) | |
Neogene | Myanmar | Emys sp. (215569) | |
Burdigalian | Switzerland | Emys sp. (69539) | |
MN 5 - MN 6 | Portugal | Emys sp. (65912) | |
Late/Upper Clarendonian | USA (Oregon) | Clemmys owyheensis (19036) | |
Hemphillian | USA (California) | Clemmys marmorata (19396 19397 19402 19407 19408) | |
early Early Hemphillian - late Early Hemphillian | USA (Idaho) | Clemmys owyheensis (218109 218111) | |
Turolian | Russian Federation (Rostovskaya Oblast') | Emys sp. (174455) | |
Early/Lower Pliocene | Moldova | Emys sp. (198590) | |
Ruscinian | China (Shanxi) | Ocadia sinensis (42273) | |
Pliocene | Slovakia | Emys orbicularis (193251) | |
Pliocene | Ukraine (Odessa) | Emys sp. (76502) | |
Pliocene - Pleistocene | Italy (Umbria) | Emys orbicularis (51684) | |
Blancan | USA (Idaho) | Clemmys owyheensis (19808) | |
MN 15 | Slovakia (Trenčín) | Emys orbicularis (34255) | |
MN 15 | Poland | Emys orbicularis, Emys wermuthi (33396) | |
MN 15 - MN 16 | Poland (Siedlce) | Emys wermuthi (34840) | |
Piacenzian | Germany (North Rhine-Westphalia) | Emys sp. (199747) | |
Piacenzian - Gelasian | USA (California) | Clemmys marmorata (166974) | |
Villanyian | Hungary (Fejer) | Emys sp., Clemmys sp. (35374) | |
Villanyian | Slovakia | Emys orbicularis (41026) | |
MN 17 | France (Haute-Loire) | Emys orbicularis (47133) | |
Early/Lower Pleistocene | Slovakia | Emys orbicularis (193253) | |
Early/Lower Pleistocene | Croatia (Istria) | Emys orbicularis (40500) | |
Early/Lower Pleistocene | Germany (Thüringen) | Emys orbicularis (43020 45063 45086) | |
Early/Lower Pleistocene | Netherlands | Emys sp. (189574) | |
Early/Lower Pleistocene | Greece (Mygdonia) | Emys orbicularis (200084) | |
Early/Lower Pleistocene | Hungary (Barany) | Emys orbicularis (35457) | |
Pleistocene | Greece (Peloponnesus) | Emys orbicularis (182721) | |
Pleistocene | Slovakia | Emys orbicularis (193252 193254 193255) | |
Pleistocene | Italy (Sardinia) | Emys obicularis (190859) | |
Pleistocene | Malta | Emys orbicularis (190860) | |
Pleistocene | United Kingdom | Emys falconeri (34978) | |
Pleistocene | France | Emys orbicularis (195262) | |
Pleistocene | United Kingdom (England) | Emys orbicularis (125089) | |
Calabrian | Spain (Palencia) | Emys orbicularis (205231) | |
Calabrian | Italy (Lombardia) | Emys europaea (190852) | |
Irvingtonian | USA (California) | Clemmys marmorata (20327) | |
Middle Pleistocene | Czech Republic | Emys orbicularis (195228) | |
Middle Pleistocene | France (Midi-Pyrenees) | Emys orbicularis (195222) | |
Middle Pleistocene | Netherlands | Emys orbicularis (109704) | |
Middle Pleistocene (interglacial) | United Kingdom (Somerset) | Emys orbicularis (195199) | |
Middle Pleistocene | Spain (Malaga) | Emys orbicularis (195564) | |
Middle Pleistocene | Greece | Emys orbicularis (194472 194474 194476 194477) | |
Middle Pleistocene | United Kingdom | Emys orbicularis (34981 195119) | |
Middle Pleistocene | United Kingdom (England) | Emys orbicularis (77754 195198) | |
Middle Pleistocene | Spain (Madrid) | Emys orbicularis (195267) | |
Middle Pleistocene | Italy (Capri Island) | Emys sp. (43016) | |
Middle Pleistocene | Greece (Peloponnesos) | Emys orbicularis (200080 200081 200082) | |
Middle Pleistocene | France (Corse) | Emys orbicularis (69062) | |
Rancholabrean | USA (California) | Clemmys marmorata (84880 84885 85314) | |
Late/Upper Pleistocene | United Kingdom | Emys orbicularis (93804 93805 93816) | |
Late/Upper Pleistocene | Italy (Sardinia) | Emys orbicularis (190854 226836) | |
Late/Upper Pleistocene | Croatia | Emys orbicularis (94487) | |
Late/Upper Pleistocene | Spain | Emys orbicularis (122309) | |
Late/Upper Pleistocene | France | Emys orbicularis (86273 195238) | |
Late/Upper Pleistocene | USA (California) | Clemmys marmorata (20273 79630 155409 190889) | |
Late/Upper Pleistocene | Germany (Brandenburg) | Emys orbicularis (198613) | |
Late/Upper Pleistocene | Monaco | Emys orbicularis (195265) | |
Late/Upper Pleistocene (interglacial) | USA (Sussex) | Emys orbicularis (195208) | |
Late/Upper Pleistocene | Portugal (Leiria) | Emys orbicularis (230380) | |
Late/Upper Pleistocene | Taiwan (N/A) | Ocadia sinensis (208971) | |
Late/Upper Pleistocene (interglacial) | United Kingdom (East Suffolk) | Emys orhicularis (195202) | |
Late/Upper Pleistocene (interglacial) | United Kingdom (Suffolk) | Emys orbicularis (195204 195210) | |
Late/Upper Pleistocene (interglacial) | United Kingdom (Norfolk) | Emys orbicularis (195206) | |
Late/Upper Pleistocene (interglacial) | USA (California) | Clemmys marmorata (20628 20642 20644) | |
Late/Upper Pleistocene - Holocene | Greece (Argolida) | Emys sp., Mauremys sp. (182709) | |
Holocene | China (Henan) | Testudo anyangensis (195288) | |
Holocene | Greece (Attica) | Emys orbicularis (183130) | |
Holocene | Bulgaria | Emys clemys (232847) | |
Holocene | Germany | Emys orbicularis (216169) | |
Holocene | Bulgaria (Tolbuchin) | Emys orbicularis (227255) | |
Holocene | Spain (Madrid) | Emys orbicularis (205224) | |
Holocene | Italy (Veneto) | Emys orbicularis (193569) |