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Champsosaurus
Taxonomy
Champsosaurus was named by Cope (1876). It is not extant. Its type is Champsosaurus profundus. It is the type genus of Champsosauridae.
It was assigned to Rhynchocephalia by Baur (1887); to Choristodera by Cope (1876), Brown (1905) and Russell (1956); to Champsosauridae by Hay (1902), Hay (1930), Estes (1964), Estes and Berberian (1970), Erickson (1972), Estes (1976), Erickson (1981), Carroll (1988), Evans (1990), Evans (1991), Cvancara and Hoganson (1993), Gao and Fox (1998), Peng et al. (2001) and Gao and Fox (2005); and to Neochoristodera by Matsumoto et al. (2007), Matsumoto and Evans (2010), Matsumoto et al. (2013) and Matsumoto et al. (2019).
It was assigned to Rhynchocephalia by Baur (1887); to Choristodera by Cope (1876), Brown (1905) and Russell (1956); to Champsosauridae by Hay (1902), Hay (1930), Estes (1964), Estes and Berberian (1970), Erickson (1972), Estes (1976), Erickson (1981), Carroll (1988), Evans (1990), Evans (1991), Cvancara and Hoganson (1993), Gao and Fox (1998), Peng et al. (2001) and Gao and Fox (2005); and to Neochoristodera by Matsumoto et al. (2007), Matsumoto and Evans (2010), Matsumoto et al. (2013) and Matsumoto et al. (2019).
Species
Synonyms
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Synonymy list
Year | Name and author |
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1876 | Champsosaurus Cope pp. 348-350 |
1876 | Champsosaurus profundus Cope pp. 350-351 |
1876 | Champsosaurus annectens Cope pp. 351-352 |
1876 | Champsosaurus brevicollis Cope pp. 352-353 |
1876 | Champsosaurus vaccinsulensis Cope p. 353 |
1881 | Champsosaurus australis Cope |
1882 | Champsosaurus saponensis Cope |
1882 | Champsosaurus puercensis Cope p. 195 |
1882 | Champsosaurus saponensis Cope p. 196 |
1887 | Champsosaurus Baur p. 100 |
1902 | Champsosaurus Hay p. 460 |
1902 | Champsosaurus annectens Hay p. 460 |
1902 | Champsosaurus australis Hay p. 461 |
1902 | Champsosaurus brevicollis Hay p. 461 |
1902 | Champsosaurus profundus Hay p. 461 |
1902 | Champsosaurus puercensis Hay p. 461 |
1902 | Champsosaurus saponensis Hay p. 461 |
1902 | Champsosaurus vaccinsulensis Hay p. 461 |
1902 | Champsosaurus annectens Lambe p. 45 |
1905 | Champsosaurus Brown |
1930 | Plesiosaurus vaccinsulensis Hay p. 116 |
1930 | Champsosaurus Hay p. 132 |
1930 | Champsosaurus annectens Hay p. 133 |
1930 | Champsosaurus australis Hay p. 133 |
1930 | Champsosaurus brevicollis Hay p. 133 |
1930 | Champsosaurus profundus Hay p. 133 |
1930 | Champsosaurus puercensis Hay p. 133 |
1930 | Champsosaurus saponensis Hay p. 133 |
1933 | Champsosaurus inflatus Parks p. 131 |
1933 | Champsosaurus inelegans Parks p. 135 |
1956 | Champsosaurus Russell p. 4 |
1964 | Champsosaurus Estes p. 99 |
1970 | Champsosaurus Estes and Berberian p. 4 |
1972 | Champsosaurus Erickson p. 8 |
1976 | Champsosaurus Estes p. 507 |
1981 | Champsosaurus Erickson |
1988 | Champsosaurus Carroll |
1990 | Champsosaurus Evans |
1991 | Champsosaurus Evans |
1993 | Champsosaurus Cvancara and Hoganson p. 15 |
1993 | Champsosaurus profundus Cvancara and Hoganson p. 15 |
1998 | Champsosaurus Gao and Fox p. 315 |
2001 | Champsosaurus Peng et al. p. 32 |
2005 | Champsosaurus Gao and Fox p. 439 |
2007 | Champsosaurus Matsumoto et al. p. 342 |
2010 | Champsosaurus Matsumoto and Evans |
2013 | Champsosaurus Matsumoto et al. |
2019 | Champsosaurus Matsumoto et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Champsosaurus Cope 1876
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†Champsosaurus albertensis Parks 1927
†Champsosaurus ambulator Brown 1905
†Champsosaurus gigas Erickson 1972
†Champsosaurus laramiensis Brown 1905
†Champsosaurus lindoei Gao and Fox 1998
†Champsosaurus natator Parks 1933
†Champsosaurus norelli Brownstein 2022
†Champsosaurus tenuis Erickson 1981
Invalid names: Champsosaurus annectens Cope 1876 [nomen vanum], Champsosaurus australis Cope 1881 [nomen vanum], Champsosaurus brevicollis Cope 1876 [nomen vanum], Champsosaurus inelegans Parks 1933 [nomen vanum], Champsosaurus inflatus Parks 1933 [nomen vanum], Champsosaurus profundus Cope 1876 [nomen vanum], Champsosaurus puercensis Cope 1882 [nomen vanum], Champsosaurus saponensis Cope 1882 [nomen vanum], Champsosaurus vaccinsulensis Cope 1876 [nomen vanum]
Diagnosis
Reference | Diagnosis | |
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K. -Q. Gao and R. C. Fox 1998 | Champsosaurus can be diagnosed by a suite of derived character states as follows: great elongation of snouth, exceeding 1/2 of skull length; interorbital width narrower than diameter of orbit; lacrimal strongly reduced to small triangle; postorbital excluded from formation of orbit; premaxilla/vomer contact lost; internarial present; choanae in posterior portion, in correspondence with elongation of vomer; interpterygoid foramen small, enclosed both anteriorly and posteriorly by pterygoids and located near posterior border of suborbital fenestra; suborbital fenestra shortened and kidney-shaped pterygoid/parasphenoid articulation fused; craniomandibular joint anterior to level of occipital condyle; neomorph forming most of ventral border of posttemporal fenestra; paraoccipital process strongly deflected ventrally; basal tubera of basisphenoid expanded posteroventrally and wing-like; elongation of mandibular symphysis to over 1/2 of length of tooth row; splenial strongly intervening in mandibular symphysis. |
Measurements
No measurements are available
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Source: g = genus, c = class, subp = subphylum | |||||
References: Carroll 1988, Bryant 1989, Hendy et al. 2009 |
Age range: base of the Late/Upper Cenomanian to the top of the Late/Upper Paleocene or 99.60000 to 55.80000 Ma
Collections (223 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Late/Upper Cenomanian | Canada (Nunavut) | C. sp. (45559) | |
Campanian | Canada (Alberta) | C. sp. (34375) | |
Judithian | USA (Wyoming) | C. natator (14458) | |
Late/Upper Campanian | USA (Montana) | C. sp. (59373) | |
Late/Upper Campanian | Canada (Alberta) | C. annectens (133004) C. natator (22695 22696 22697 22698 34302 34303 34305 34307 34309 34310 34311 34312 34313 34315 34319 34320 34321 34322 45238 45260 45261 45262 45264 47784 47793 47807 47811 47812 47814 47816 189515) C. sp. (22659 36264 45671 45674 45822 45892 45893 45897 45898 98674 98677 219724 227125) | |
Middle Campanian | Canada (Alberta) | C. lindoei (117922 118188 118190) C. natator (45240 47794 47795 47809 117923 117927 117928 117929 117930) C. sp. (45560 45561 45564 45565 45566 45567 45670 45820 118189 181024) | |
Middle Campanian | USA (Montana) | C. sp. (14437 22691) | |
Middle Campanian - Late/Upper Campanian | Canada (Saskatchewan) | C. sp. (14494 75030 229975) | |
Middle Campanian - Late/Upper Campanian | Canada (Alberta) | C. annectens (150749) C. sp. (45819 45821 45894 62722) | |
Middle Campanian - Late/Upper Campanian | USA (Montana) | C. lindoei (231393) C. sp. (14490 50907 59217) | |
Late/Upper Campanian - Early/Lower Maastrichtian | Canada (Alberta) | C. albertensis (117924) C. sp. (34373 52072 52249 163050) | |
Maastrichtian | USA (South Dakota) | C. sp. (45391) | |
Maastrichtian | USA (North Dakota) | C. sp. (45135 45380) | |
Early/Lower Maastrichtian | Canada (Alberta) | C. sp. (14502 52468 55363 135740) | |
Lancian | USA (Colorado) | C. sp. (14640) | |
Lancian | USA (Montana) | C. natator (14512 14533 14552 14615) C. sp. (14508 14547 14548 14560 14580 14613 14653) C. sp., C. natator (14549 14582 14639 14654) | |
Lancian | Canada (Saskatchewan) | C. sp. (14655) | |
Lancian | USA (Wyoming) | C. sp. (14505 14514 14519 14524 14538 14540 14557 14566 75447) | |
Lancian | USA (South Dakota) | C. sp. (14537 192251) | |
Lancian | USA (North Dakota) | C. laramiensis (14620 14621) C. sp. (14610 137601 137704 137706 137707 137710 137711 137712 137713 137715 137716 137724 137728 137731) | |
Late/Upper Maastrichtian | Canada (Alberta) | C. sp. (14569) | |
Late/Upper Maastrichtian | USA (Wyoming) | C. sp. (14585) | |
Late/Upper Maastrichtian | USA (Montana) | C. laramiensis, C. ambulator (131865) C. sp. (13142 14674 88990 183109 183110) | |
Late/Upper Maastrichtian | USA (South Dakota) | C. sp. (45388 45389 45390 48551 50082 150891 150892 150893 150894 150895 224484) | |
Late/Upper Maastrichtian | USA (North Dakota) | C. laramiensis (231465) C. sp. (45099 45102 45109 45110 45111 45112 45114 45117 45118 45120 45122 45124 45130 45134 45136 45144 45145 45299 45381 45382 57919 137720 141583) | |
Late/Upper Maastrichtian | Canada (Saskatchewan) | C. laramiensis (180638) C. sp. (14555 35251 45675 54105) | |
Puercan | USA (Utah) | C. sp. (14692 14790) | |
Puercan | USA (Colorado) | C. sp. (176747 205944 206079 209904) | |
Puercan | USA (Wyoming) | C. sp. (14729 14730 14796 14797 14805) | |
Puercan | USA (Montana) | C. sp. (14766 123518) | |
Danian | USA (North Dakota) | C. sp. (14962) | |
Torrejonian | USA (Montana) | C. sp. (14874) | |
Torrejonian | USA (Texas) | C. sp. (110177) | |
Torrejonian | USA (Wyoming) | C. sp. (14912) | |
Ti3 | Canada (Alberta) | C. sp. (14992) | |
Ti4 | Canada (Saskatchewan) | C. gigas (14978) | |
Tiffanian | USA (North Dakota) | C. gigas (27640) C. sp. (14931) C. sp., C. gigas (15000) C. tenuis (118170) | |
Tiffanian | USA (Wyoming) | C. norelli (15029) C. sp. (15017) | |
Late/Upper Paleocene | France (Champagne-Ardenne) | C. sp. (117931) | |
Late/Upper Paleocene | France (Oise) | C. sp. (166598) | |
Late/Upper Paleocene | USA (North Dakota) | C. gigas (78394) | |
Cf2 | USA (Wyoming) | C. sp. (46679) | |
Cf3 | USA (Wyoming) | C. sp. (46676 46678) | |
Clarkforkian | USA (Wyoming) | C. gigas (15181 15212) | |
Clarkforkian | USA (Montana) | C. sp. (137834) |