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Vadaravis brownae

Reptilia - Pelecaniformes - Threskiornithidae

Taxonomy
Vadaravis brownae was named by Smith et al. (2013) [Vadaravis brownae possesses the following autapomorphies, which are unique among living birds: (1) two (cranial and caudal) small and discrete pneumatopores on the lateral sides of the two caudal-most thoracic centra; and (2) a caudoventrally located pisiform process of the carpometacarpus that projects primarily ventrally and only weakly cranially. In addition to these autapomorphies, Vadaravis brownae can also be diagnosed by the following unique combination of characters: (1) between 12 and 13 synsacral vertebrae; (2) transverse processes of caudal-most synsacral vertebra paddle-shaped in ventral aspect; (3) eight caudal vertebrae (counting the pygostyle as a single element); (4) transverse processes of the caudal vertebrae abruptly disappear in the last free vertebra before the pygostyle; (5) absence of uncinate processes on the thoracic ribs; (6) pneumatic foramina present at the cranial end of the dorsal surface of the sternum, divided medially by a lamina; (7) well-developed ‘balland-socket’ joint between the scapula and coracoid; (8) a spina dorsolateralis ilii that does not extend quite as far caudally as the terminal end of the ischium; and (9) a pons supratendineus that is located in the middle of the distal tibiotarsus. Because Vadaravis is similar in overall size and limb proportions, and known from the same deposits as Presbyornis (Ericson, 2000), a differential diagnosis between the two taxa is warranted, and may aid in the future identification and referral of specimens to Vadaravis. FMNH PA 718 can be distinguished from Presbyornis pervetus Wetmore, 1926, by the following features: (1) posterior thoracic vertebrae less elongate craniocaudally, with two rather than a single pneumatopore; (2) fewer synsacral vertebrae cranial to the cranial end of fossa renalis (section I of Boas, 1934); (3) presence of a recessus caudalis fossa in the ilium; (4) sternal end of coracoid less elongate mediolaterally, and straight rather than weakly concave caudally; (5) coracoid shaft more elongate; (6) tuberculum coracoideum more ventrally placed on proximal scapula; (7) deeper incisure between trabecula lateralis and trabecula intermedia of the sternum (Livezey, 1997); (8) shorter symphysis intermetacarpalis; (9) absence of a preacetabular tuberculum; (10) fusion between the ilia and synsacrum; (11) femur more robust; (12) crista cnemialis lateralis of tibiotarsus convex proximally rather than straight (Ericson, 2000); and (13) distinct distolateral hook in lateral end of crista cnemialis lateralis of tibiotarsus (Ericson, 2000). Vadaravis can be distinguished from the early–middle Eocene stem-threskiornithid Rhynchaeites messelensis Wittich, 1898 (see also Peters, 1983; Mayr, 2002; Mayr and Bertelli, 2011), by the following features: (1) absence of a notarium; (2) facies articularis clavicularis of the coracoid not strongly everted medially; (3) presence of trabecula intermedia on the caudal edge of the sternum; (4) shorter symphysis intermetacarpalis; and (5) much larger size.]. Its type specimen is FMNH PA 718, a partial skeleton (a partial postcranial skeleton including portions of the axial column and associated ribs, sternum, furcula, scapula, coracoid, both humeri, both ulnae, radius,), and it is a 3D body fossil. Its type locality is Fossil Lake Sample Site H-1, which is in a Wasatchian lacustrine - large horizon in the Green River Formation of Wyoming.

Synonymy list
YearName and author
2013Vadaravis brownae Smith et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
Archosauromorpha(Huene 1946)
Crocopoda
RankNameAuthor
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Theropoda()
Neotheropoda
AverostraPaul 2002
Tetanurae
Coelurosauria()
Maniraptora
Paraves
classAves
subclassNeornithesGadow 1893
Aequornithes
Feraequornithes
superorderPelecanimorphaeHuxley 1867
orderPelecaniformesSharpe 1891
familyThreskiornithidaeRichmond 1917
genusVadaravis
speciesbrownae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Vadaravis brownae Smith et al. 2013
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Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yeso
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: solitaryo
Environment: terrestrialf
Locomotion: actively mobileo
Life habit: volantf
Diet: carnivoref
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2009-01-03 17:09:47
Modified: 2009-01-03 19:09:47
Source: f = family, o = order
References: Marsh 1875, Terborgh et al. 1990

Age range: Wasatchian or 54.90000 to 50.50000 Ma

Collections: one only


Time interval Ma Country or state Original ID and collection number
Wasatchian54.9 - 50.5USA (Wyoming) Vadaravis brownae (type locality: 74617)