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Scalpellomorpha

Thecostraca - Scalpellomorpha

Taxonomy
Scalpellomorpha was named by Newman (1987). It is extant.

It was reranked as the order Scalpellomorpha by Chan et al. (2021).

It was assigned to Pedunculata by Carriol and Collins (2000); to Scalpelliformes by Jones (2007); and to Thoracicalcarea by Chan et al. (2021).

Synonymy list
YearName and author
1987Scalpellomorpha Newman
2000Scalpellomorpha Carriol and Collins p. 142
2007Scalpellomorpha Jones p. 291
2021Scalpellomorpha Chan et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
RankNameAuthor
subphylumCrustaceaBrünnich 1772
classThecostraca(Gruvel 1905)
subclassCirripedia(Rafinesque 1816)
infraclassThoracica(Darwin 1854)
superorderThoracicalcarea()
orderScalpellomorpha(Newman 1987)
orderScalpellomorpha(Newman 1987)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Scalpellomorpha Newman 1987
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Superfm. Lepadoidea Darwin 1852
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Fm. Heteralepadidae Nilsson-Cantrell 1921
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G. Alepas Rang 1829
G. Heteralepas Pilsbry 1907
G. Koleolepas Stebbing 1900
G. Paralepas Pilsbry 1907
Fm. Lepadidae Darwin 1851
+
G. Conchoderma von Olfers 1814
G. Dosima Gray 1925
G. Hyalolepas Annandale 1906
G. Lepas Linnaeus 1758
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Subg. Lepas (Anatifa) Bruguière 1789
Subg. Lepas (Lepas) Linnaeus 1758
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†Lepas (Lepas) clifdenica Buckeridge 1983
†Lepas (Lepas) hyperumbonica Buckeridge 1983
Lepas anatifera Linnaeus 1758
†Lepas sattmanni Harzhauser and Schloegl 2012
G. †Pristinolepas Buckeridge 1983
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Fm. Malacolepadidae Hiro 1937
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G. Arcalepas Jones and Morton 2009
G. Malacolepas Hiro 1933
Fm. Poecilasmatidae Annandale 1909
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G. †Archoxynaspis Van Syoc and Dekelboum 2011
G. Dianajonesia Kocak and Kemal 2008
G. Dichelaspis Darwin 1852
G. Glyptelasma Pilsbry 1907
G. Megalasma Hoek 1883
G. Microlepas Hoek 1907
G. Minyaspis Van Syoc and Dekelboum 2011
G. Octolasmis Gray 1825
G. Oxynaspis Darwin 1852
G. Pagurolepas Stubbings 1940
G. Poecilasma Darwin 1851
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†Poecilasma miocaenica Reuss 1864
G. Rugilepas Grygier and Newman 1991
G. Scleraspis Van Syoc and Dekelboum 2012
G. Temnaspis Fischer 1884
G. Trilasmis Hinds 1844
Invalid names: Microlepadidae Zevina 1980 [synonym], Oxynaspididae Gruvel 1905 [synonym]
Fm. Rhizolepadidae Zevina 1980
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G. Rhizolepas Gay 1939
Superfm. Neolepadoidea Yamaguchi et al. 2004
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Fm. Neobrachylepadidae Newman and Yamaguchi 1995
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G. Neobrachylepas Newman and Yamaguchi 1995
Fm. Neolepadidae Yamaguchi et al. 2004
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G. Ashinkailepas Yamamoto et al. 2004
G. Leucolepas Southward and Jones 2003
G. Neolepas Newman 1979
G. †Stipilepas Carriol 2016
G. Vulcanolepas Southward and Jones 2003
Fm. Neoverrucidae Newman 1989
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G. Imbricaverruca Newman 2000
G. Neoverruca Newman and Yamaguchi 1995
Superfm. Scalpelloidea Pilsbry 1907
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Fm. †Proverrucidae Newman 1989
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G. †Proverruca Withers 1914
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†Proverruca dentifer Gale 2014
Fm. Scalpellidae Pilsbry 1907
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Subfm. Amigdoscalpellinae Gale 2016
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G. Amigdoscalpellum Zevina 1978
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G. Catherinum Zevina 1978
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†Catherinum anglicum Gale 2016
G. Weltnerium Zevina 1978
G. Anguloscalpellum Zevina 1978
G. Barbascalpellum Zevina 1978
Subfm. Brochiinae Zevina 1978
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G. Australscalpellum Newman and Ross 1971
G. Brochia Newman and Ross 1971
G. Compressoscalpellum Zevina 1978
G. Diceroscalpellum Zevina 1978
Subfm. Meroscalpellinae Zevina 1978
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G. Abathescalpellum Newman and Ross 1971
G. Alcockianum Zevina 1978
G. Annandaleum Newman and Ross 1978
G. Gymnoscalpellum Newman and Ross 1971
G. Hamatoscalpellum Zevina 1978
G. Litoscalpellum Newman and Ross 1971
G. Meroscalpellum Zevina 1978
G. Neoscalpellum Pilsbry 1907
G. Pilsbryiscalpellum Zevina 1978
G. Planoscalpellum Zevina 1978
G. Pteroscalpellum Zevina 1978
Subfm. Scalpellinae Pilsbry 1907
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G. Arcoscalpellum Hoek 1907
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†Arcoscalpellum bakeri Collins 1973
†Arcoscalpellum campus Collins 1973
†Arcoscalpellum choctawensis Weisbord 1977
†Arcoscalpellum euglyphum Withers 1924
†Arcoscalpellum gaultinum Withers 1935
†Arcoscalpellum habanense Withers 1953
†Arcoscalpellum hartleyi Withers 1936
†Arcoscalpellum hubrichti Collins 1973
†Arcoscalpellum moskvini Alekseev 2009
†Arcoscalpellum pulchellum Bosquet 1854
†Arcoscalpellum renevieri Mayer and Gumbel 1861
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Invalid names: Arcoscalpellum loczyi Szorenyi 1934 [synonym]
†Arcoscalpellum singletoni Withers 1953
†Arcoscalpellum toulmini Weisbord 1977
†Arcoscalpellum withersi Collins 1973
G. †Arcuatoscalpellum Gale 2016
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G. Diotascalpellum Gale 2016
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†Diotascalpellum acies Gale 2020
†Diotascalpellum fossula Darwin 1851
†Diotascalpellum gracile Bosquet 1854
G. Graviscalpellum Foster 1980
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†Graviscalpellum chapmani Withers 1924
†Graviscalpellum crassiforme Buckeridge 1983
†Graviscalpellum dunni Chapman and Crespin 1928
†Graviscalpellum euglyphum Withers 1924
†Graviscalpellum grantmackiei Buckeridge 1983
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Invalid names: Arcoscalpellum distinctum Grant-Mackie 1965 [replaced]
†Graviscalpellum ungulatum Withers 1913
G. †Jaegerscalpellum Gale 2019
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†Jaegerscalpellum comptum Withers 1910
†Jaegerscalpellum politum Darwin 1851
G. Regioscalpellum Gale 2016
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†Regioscalpellum maximum Sowerby 1829
†Regioscalpellum sanchezae Withers 1926
G. Scalpellum Leach 1818
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†Scalpellum accumulatum Withers 1910
†Scalpellum aduncatum Withers 1910
Scalpellum australe Buckeridge 1983
†Scalpellum cyphum Withers 1910
†Scalpellum dalpiazi Withers 1953
†Scalpellum dissimile Withers 1910
†Scalpellum gibbum Pilsbry 1916
†Scalpellum lineatum Darwin 1851
†Scalpellum longissimum Withers 1911
†Scalpellum paratethyianum Harzhauser and Schloegl 2012
†Scalpellum parvulum Withers 1914
†Scalpellum pfeifferi Weithofer 1887
Scalpellum scalpellum Linnaeus 1758
†Scalpellum subquadratum Meyer and Aldrich 1886
G. Zevinaella Shalava and Newman 2015
Invalid names: Arcoscalpellinae Zevina 1978 [synonym]
Subfm. Scalpellopsinae Zevina 1978
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G. Scalpellopsis Broch 1921
G. Sinoscalpellum Ren and Sha 2014
G. Teloscalpellum Zevina 1978
G. Trianguloscalpellum Zevina 1978
G. Vertebroscalpellum Newman and Ross 1998
Subfm. †Virgiscalpellinae Gale 2020
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G. †Collinslepas Gale 2020
G. †Virgilepas Gale 2020
G. †Virgiscalpellum Withers 1935
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†Virgiscalpellum beisseli Bosquet and Müller 1857
†Virgiscalpellum gabbi Pilsbry 1933
†Virgiscalpellum inornatum Zullo and Sohl 1985
†Virgiscalpellum wrighti Withers 1946
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Locomotion: stationaryc
Attached: yesc
Epibiont: yesc
Life habit: epifaunalc
Diet: suspension feederc
Vision: limitedc
Created: 2008-03-04 07:51:33
Modified: 2011-06-14 02:28:17
Source: c = class
Reference: Kiessling 2004

Age range

Maximum range based only on fossils: base of the Early/Lower Hauterivian to the top of the Zanclean or 132.60000 to 3.60000 Ma
Minimum age of oldest fossil (stem group age): 125.77 Ma

Collections (158 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Hauterivian132.6 - 125.77Germany Jaegerscalpellum elegans (91586)
Early/Lower Hauterivian132.6 - 125.77France Scalpellidae indet. (59653)
Early/Lower Aptian121.4 - 119.57United Kingdom (England) Arcoscalpellum comptum, Virgiscalpellum wrighti (171903)
Aptian121.4 - 113.2United Kingdom (England) Virgiscalpellum wrighti (1747)
Late/Upper Aptian119.57 - 113.2United Kingdom Scalpellum comptum (211496)
Early/Lower Albian113.2 - 110.1United Kingdom (Kent) Scalpellum accumulatum (211495)
Albian113.2 - 100.5France Arcoscalpellum arcuatum (160155) Arcoscalpellum arcuatum, Arcoscalpellum comptum (160148)
Albian113.2 - 100.5United Kingdom (England) Scalpellum arcuatum (86975 211493)
Middle Albian110.1 - 106.3United Kingdom (England) Arcoscalpellum arcuatum (211460 211462) Arcoscalpellum arcuatum, Arcoscalpellum comptum (211447 211448 211449 211454) Arcoscalpellum comptum (211455 211459)
Middle Albian110.1 - 106.3USA (Texas) Virgiscalpellum marysensis (212388 212399)
Late/Upper Albian106.3 - 100.5United Kingdom (England) Arcoscalpellum arcuatum (211461 211470 211474 211475 211476 211478) Arcoscalpellum arcuatum, Arcoscalpellum comptum (211450 211457 211463 211468 211469 211471) Arcoscalpellum arcuatum, Arcoscalpellum comptum, Arcoscalpellum angustatum (211452 211458 211464 211472 211473) Arcoscalpellum arcuatum, Arcoscalpellum trilineatum, Arcoscalpellum gaultinum (211465) Arcoscalpellum lineatum, Arcoscalpellum arcuatum, Arcoscalpellum comptum, Arcoscalpellum angustatum (211451) Jaegerscalpellum politum, Arcuatoscalpellum arcuatum (210476)
Late/Upper Albian106.3 - 100.5USA (Texas) Diotascalpellum acies (212380 212381 212390)
Late/Upper Albian106.3 - 100.5USA (Oklahoma) Virgiscalpellum rugosum (212382 212386 212389)
Cenomanian100.5 - 93.9USA (Texas) Scalpellum sp. (6492)
Early/Lower Cenomanian100.5 - 93.9USA (Texas) Arcuatoscalpellum wacoensis, Virgilepas sp. (212394)
Early/Lower Cenomanian100.5 - 93.9United Kingdom (England) Scalpellum parvulum (160379 211487 211488)
Late/Upper Cenomanian100.5 - 93.9USA (Arizona) Scalpellum sp. (53488 53489 53492)
Middle Cenomanian100.5 - 93.9United Kingdom (England) Scalpellum aduncatum (211498)
Middle Cenomanian - Late/Upper Cenomanian100.5 - 93.9United Kingdom (England) Scalpellum cyphum (211497) Scalpellum trilineatum (211492)
Early/Lower Turonian93.9 - 89.8Czechia Scalpellidae indet. (90516 90518) Scalpellum sp. (146401 146500 146501)
Late/Upper Turonian93.9 - 89.8Kazakhstan Arcoscalpellum angustatum (210443)
Late/Upper Turonian93.9 - 89.8Czechia Diotascalpellum angustatum (210303 210305 210307 210308) Regioscalpellum maximum (210304)
Middle Turonian93.9 - 89.8United Kingdom (England) Scalpellum dissimile (211499)
Santonian85.7 - 83.6USA (Kansas) Scalpellidae indet. (94652)
Late/Upper Santonian - Early/Lower Campanian85.7 - 72.2Russian Federation (Penza) Arcoscalpellum moskvini (210370)
Campanian83.6 - 72.2USA (North Carolina) Virgiscalpellum inornatum (7613)
Early/Lower Campanian83.6 - 72.2United Kingdom (England) Scalpellum darwinianum, Scalpellum longissimum (159324)
Early/Lower Campanian83.6 - 72.2Sweden (Skane) Arcoscalpellum sp., Poecilasma sp., Virgiscalpellum sp., Arcoscalpellum maximum, Arcoscalpellum scaniensis, Proverruca dentifer (210592)
Early/Lower Campanian83.6 - 72.2Poland Arcoscalpellinae indet. (212511) Virgiscalpellum sp. (212514) Virgiscalpellum sp., Arcoscalpellinae indet., Arcoscalpellum fossula (212513)
Early/Lower Campanian83.6 - 72.2USA (Alabama) Arcoscalpellum hubrichti (172589 172617 203110 203111 210612 210613 210614 210615 210616 210621)
Early/Lower Campanian83.6 - 72.2USA (Mississippi) Arcoscalpellum hubrichti (210609)
Early/Lower Campanian83.6 - 72.2France Virgiscalpellum beisseli, Arcoscalpellum fossula (59776)
Late/Upper Campanian83.6 - 72.2USA (Tennessee) Scalpellum sp. (88357)
Late/Upper Campanian83.6 - 72.2United Kingdom (England) Diotascalpellum fossula, Catherinum anglicum (157759) Regioscalpellum maximum, Catherinum anglicum (210319)
Late/Upper Campanian83.6 - 72.2Kazakhstan Virgiscalpellum mangyshlakense (210371)
Late/Upper Campanian83.6 - 72.2Belgium Arcoscalpellum fossula (211444) Arcoscalpellum maximum (211443) Arcoscalpellum maximum, Virgiscalpellum beisseli, Arcoscalpellum fossula (211442)
Late/Upper Campanian83.6 - 72.2USA (Delaware) Arcoscalpellum bakeri, Arcoscalpellum withersi (83133)
Late/Upper Campanian83.6 - 72.2Austria Arcoscalpellum maximum (48008)
Middle Campanian83.6 - 72.2United Kingdom (England) Scalpellum darwinianum (211481 211482)
Middle Campanian83.6 - 72.2USA (Mississippi) Virgiscalpellum sp. (172903 210599)
Middle Campanian83.6 - 72.2Poland Arcoscalpellinae indet., Arcoscalpellum fossula (212521 212525) Arcoscalpellum fossula (212520 212526) Arcoscalpellum sp., Virgiscalpellum sp., Arcoscalpellinae indet. (212524) Virgiscalpellum sp., Arcoscalpellum fossula (212522 212523)
Campanian - Early/Lower Maastrichtian83.6 - 66.0Poland Arcoscalpellum maximum, Arcoscalpellum fossula (212527)
Late/Upper Campanian - Early/Lower Maastrichtian83.6 - 66.0USA (Mississippi) Arcoscalpellum bakeri, Arcoscalpellum campus, Arcoscalpellum withersi (210594) Arcoscalpellum campus (210600) Virgiscalpellum sp. (210596)
Early/Lower Maastrichtian72.2 - 66.0Kazakhstan Arcoscalpellum maximum (210449) Arcoscalpellum moskvini (210368 210369)
Early/Lower Maastrichtian72.2 - 66.0USA (Mississippi) Arcoscalpellum bakeri, Arcoscalpellum campus, Arcoscalpellum withersi (172643 210602 210603 210604 210608) Arcoscalpellum bakeri, Arcoscalpellum campus, Arcoscalpellum withersi, Virgiscalpellum gabbi (210595) Arcoscalpellum bakeri, Arcoscalpellum withersi (210601) Arcoscalpellum campus, Arcoscalpellum withersi, Virgiscalpellum gabbi (210605)
Late/Upper Maastrichtian72.2 - 66.0USA (Mississippi) Arcoscalpellum campus, Arcoscalpellum withersi (210607) Arcoscalpellum withersi (210598) Virgiscalpellum gabbi (210597)
Late/Upper Maastrichtian72.2 - 66.0Netherlands Virgiscalpellum darwinianum, Virgiscalpellum hagenowianum, Arcoscalpellum gracile, Arcoscalpellum pulchellum (138910)
Late/Upper Maastrichtian72.2 - 66.0Belgium Virgiscalpellum hagenowianum, Arcoscalpellum maximum (211446) Virgiscalpellum hagenowianum, Arcoscalpellum maximum, Arcoscalpellum fossula (211445)
Late/Upper Maastrichtian72.2 - 66.0USA (Alabama) Arcoscalpellum withersi (210620) Arcoscalpellum withersi, Virgiscalpellum gabbi (210619)
Late/Upper Maastrichtian72.2 - 66.0Kazakhstan Arcoscalpellum maximum (210446 210447 210448)
Danian66.0 - 61.66Denmark Scalpellum steenstrupi (89866)
Danian66.0 - 61.66USA (Alabama) Arcoscalpellum toulmini (2855)
Danian66.0 - 61.66Egypt Scalpellum curvatum (86950)
Thanetian59.24 - 56.0USA (New Jersey) Scalpellum conradi (3113 124966)
Ypresian56.0 - 48.07United Kingdom (England) Arcoscalpellum quadratum (76601)
Bartonian41.03 - 37.71Spain Arcoscalpellum sp. (141924)
Priabonian37.71 - 33.9USA (Louisiana) Arcoscalpellum jacksonensis (32925)
Priabonian37.71 - 33.9USA (Alabama) Oxynaspis sp. (208482)
Rupelian33.9 - 27.3Austria Arcoscalpellum renevieri (160670)
Rupelian33.9 - 27.3Hungary Arcoscalpellum renevieri (160671)
Rupelian33.9 - 27.3USA (South Carolina) Arcoscalpellum sp. (66190)
Chattian27.3 - 23.04USA (Washington) Arcoscalpellum knapptonensis, Arcoscalpellum raricostatum (160141) Arcoscalpellum sp. (91624)
Aquitanian23.04 - 20.45Austria Lepas sattmanni (99160)
Burdigalian20.45 - 15.98Germany Scalpellidae indet. (188985)
Burdigalian20.45 - 15.98Slovakia Scalpellum paratethyianum (160736)
Burdigalian20.45 - 15.98Austria Scalpellum pfeifferi (160672 160673)
Langhian15.98 - 13.82Austria Poecilasma miocaenica (160737)
Middle Miocene15.98 - 11.63Poland Lepas miocoenicus (70539)
Late/Upper Miocene11.63 - 5.333USA (Florida) Scalpellum gibbum (91537)
Zanclean5.333 - 3.6Holy See (Vatican City State) Anatifa scolpellum (209253)