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Canis armbrusteri

Osteichthyes - Carnivora - Canidae

Taxonomy
Canis armbrusteri was named by Gidley (1913). Its type specimen is USNM 7662, a mandible, and it is a 3D body fossil. Its type locality is Cumberland Cave, which is in an Irvingtonian cave horizon in Maryland.

It was synonymized subjectively with Canis lupus by Martin (1974), Miller (1976); it was recombined as Aenocyon armbrusteri by Eshelman et al. (2025).

Synonymy list
YearName and author
1913Canis armbrusteri Gidley
1979Canis armbrusteri Nowak
1980Canis armbrusteri Kurten and Anderson p. 169
2008Canis armbrusteri Wang et al.
2009Canis armbrusteri Tedford et al. p. 137 figs. 40, 43, 44, 52, 53A–D, 54A–F, 55A–D; appendices 2–4
2019Canis armbrusteri Albright et al. p. 173
2025Aenocyon armbrusteri Eshelman et al. p. 117

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
RankNameAuthor
Mammaliaformes
classMammalia
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Ferae()
Pancarnivora
CarnivoramorphaWyss and Flynn 1993
CarnivoraformesFlynn et al.
orderCarnivora
familyCanidae
subfamilyCaninaeGill 1872
genusCanis
speciesarmbrusteri

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Canis armbrusteri Gidley 1913
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Diagnosis
ReferenceDiagnosis
R. H. Tedford et al. 2009Canis armbrusteri shares with C. lupus and C. dirus the following synapomorphies: incisive foramina extend posterior to limit of canine alveoli, strong posterior expansion of paroccipital process, posterior expansion of frontal sinus to frontoparietal suture, and very reduced M1 parastyle lacking union with prepara- crista. It differs from C. lupus in that it retains premolars in which P3 and p2–p3 usually have posterior cusps, and the second posterior cusp of p4 is separate from the posterior cingulum. Like C. dirus, it also differs from C. lupus in having short-crowned and relatively straight canines, the metaconid of m1–m2 reduced, and a reduced P4 protocone. Although a sister taxon to C. dirus, C. armbrusteri lacks the other hyper- carnivorous dental features characteristic of C. dirus including a primitively retained labial cingulum on M1.
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: carnivoref
Diet 2: omnivoref
Reproduction: viviparoussubc
Created: 2005-06-08 10:11:09
Modified: 2005-08-22 19:28:06
Source: f = family, subc = subclass, c = class, subp = subphylum
References: Nowak 1991, Hendy et al. 2009, Carroll 1988, Lillegraven 1979, Ji et al. 2002

Age range: base of the Irvingtonian to the top of the Late/Upper Pleistocene or 1.40000 to 0.01170 Ma

Collections (18 total)


Time interval Ma Country or state Original ID and collection number
Calabrian1.8 - 0.774USA (Virginia) Canis armbrusteri (104698)
Calabrian1.8 - 0.774USA (Florida) Canis armbrusteri (20400 20401)
Irvingtonian1.4 - 0.21USA (Texas) Aenocyon sp. (20530) Canis dirus (20498) Canis lupus (20342)
Irvingtonian1.4 - 0.21USA (Maryland) Canis armbrusteri (type locality: 20302)
Irvingtonian1.4 - 0.21USA (Kansas) Canis armbrusteri (20537)
Irvingtonian1.4 - 0.21USA (Arizona) Canis nubilus (19617)
Irvingtonian1.4 - 0.21USA (Pennsylvania) Canis priscolatrans (20479)
Irvingtonian1.4 - 0.21USA (Florida) Canis armbrusteri (20351 20415) Canis lupus (20289)
Irvingtonian1.4 - 0.21USA (New Mexico) Canis armbrusteri (20558)
Irvingtonian1.4 - 0.21USA (Arkansas) Canis occidentalis (20290)
Irvingtonian1.4 - 0.21USA (South Carolina) Canis dirus (79919)
Late/Upper Pleistocene0.129 - 0.0117USA (South Carolina) Canis armbrusteri (152904)
Late/Upper Pleistocene (interglacial)0.129 - 0.0117USA (Florida) Canis lupus (20593)