PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Scyphozoa (jellyfish)

Scyphozoa

Taxonomy

Scyphozoa was named by Goette (1887). It is extant. It was considered polyphyletic by Kayal et al. (2013).

It was assigned to Cnidaria by Barnes (1998), Sepkoski (2002), Pacheco et al. (2015); and to Medusozoa by Kayal et al. (2013), Van Iten et al. (2014), Van Iten et al. (2016), Miller et al. (2022).

Synonymy list

Year	Name and author
1887	Scyphozoa Goette
1998	Scyphozoa Barnes p. 185
2002	Scyphozoa Sepkoski
2013	Scyphozoa Kayal et al.
2014	Scyphozoa Van Iten et al. p. 678
2015	Scyphozoa Pacheco et al. p. 14
2016	Scyphozoa Van Iten et al.
2022	Scyphozoa Miller et al. p. 2376

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Rank	Name	Author
kingdom	Animalia	()
phylum	Cnidaria	Hatschek 1888
subphylum	Medusozoa

Rank	Name	Author
class	Scyphozoa	Goette 1887
class	Scyphozoa	Goette 1887

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Cl. Scyphozoa Goette 1887 [jellyfish]

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G. †Adesmoconularia

Or. †Anabaritida

Fm. †Anabaritidae Missarzhevsky 1974

G. †Aculeochrea Val'kov and Sysoev 1970

†Aculeochrea ornata Val'kov and Sysoev 1970

†Aculeochrea rugosa Val'kov and Sysoev 1970

Invalid names: Longiochrea Val'kov and Sysoev 1970 [synonym]

G. †Anabarites Voronova and Missarzhevsky 1969

†Anabarites biplicatus Missarzhevsky 1989

Invalid names: Anabarites bisulcatus Federov 1990 [synonym]

†Anabarites compositus Rozanov et al. 1969

†Anabarites hariolus Val'kov 1987

†Anabarites hexasulcatus Missarzhevsky 1974

Invalid names: Anabaritellus xishuiensis Qian 1977 [synonym]

†Anabarites kelleri Missarzhevsky 1989

†Anabarites korobovi Rozanov and Missarzhevsky 1966

†Anabarites latus Val'kov and Sysoev 1970

Invalid names: Angustiochrea magna Bokova 1990 [synonym]

†Anabarites modestus Bokova 1985

†Anabarites natellus Val'kov and Sysoev 1970

Invalid names: Lobiochrea formosa Bokova 1990 [synonym], Lobiochrea trialata Val'kov 1989 [synonym]

†Anabarites rectus Vasil'eva 1984

†Anabarites sexalox Bengtson et al. 1990

†Anabarites signatus Missarzhevsky and Mambetov 1981

†Anabarites tristichus Rozanov et al. 1969

Invalid names: Anabarites gracilis Chen 1984 [synonym], Jakutiochrea lenta Val'kov 1987 [synonym], Jakutiochrea portentosa Val'kov 1987 [synonym], Jakutiochrea solita Val'kov 1987 [synonym]

†Anabarites trisulcatus Voronova and Missarzhevsky 1969

Invalid names: Anabarites grandis Val'kov 1982 [synonym], Anabarites obliquasulcatus Qian 1978 [synonym], Anabarites primitivus Qian and Jiang 1980 [synonym], Anabarites rotundum Qian 1977 [synonym], Anabarites rotundus Qian 1989 [synonym], Anabarites sulcoconvex Qian 1978 [synonym], Anabarites tenuistriatus Qian 1989 [synonym], Anabarites undulatus Qian 1978 [synonym]

†Anabarites trymatus Bengtson et al. 1990

†Anabarites valkovi Bokova 1990

†Anabarites voluta Rozanov et al. 1969

Invalid names: Anabaritellus Missarzhevsky 1974 [synonym], Angustiochrea Val'kov and Sysoev 1970 [synonym], Jakutiochrea Val'kov and Sysoev 1970 [synonym], Kotuites Missarzhevsky 1989 [synonym], Kugdatheca Rozanov et al. 1969 [synonym], Lobiochrea Val'kov and Sysoev 1970 [synonym], Sexangulatus Federov 1990 [synonym], Udzhaites Vasil’eva 1986 [synonym]

G. †Cambrotubulus Rozanov et al. 1969

†Cambrotubulus corniformis Elicki 1994

†Cambrotubulus decurvatus Rozanov et al. 1969

Invalid names: Cambrotubulus conicus Missarzhevsky 1989 [synonym], Cambrotubulus crassus Federov 1990 [synonym], Cambrotubulus plicativus Val'kov 1987 [synonym], Platysolenites sibirica Val'kov 1968 [synonym]

G. †Mariochrea Val'kov 1982

†Mariochrea sinuosa Val'kov 1982

Invalid names: Gastreochrea viva Val'kov 1982 [synonym]

Invalid names: Gastreochrea Val'kov 1982 [synonym]

G. †Selindeochrea Val'kov 1982

†Selindeochrea missarzhevskyi Vasil’eva 1986

†Selindeochrea tecta Val'kov 1982

†Selindeochrea ternaria Rozanov et al. 1969

†Selindeochrea tricarinata Rozanov et al. 1969

†Selindeochrea tripartita Rozanov et al. 1969

Invalid names: Anabarites tribaculatus Bokova 1990 [synonym]

G. †Tiksitheca Rozanov et al. 1969

†Tiksitheca fangxianensis Duan 1984

†Tiksitheca huangshandongensis Qian et al. 1979

†Tiksitheca licis Rozanov et al. 1969

Invalid names: Anabaritinae Missarzhevsky 1974 [empty]

G. †Australoconularia

G. †Beltanella Sprigg 1947

†Beltanella zimimica Bekker 1992

G. †Bjarmia Grazhdankin 2016

†Bjarmia cycloplerusa Grazhdankin 2016

G. †Circonularia

G. †Conularina

Or. †Coronatida

Fm. Collaspididae Haeckel 1880

G. †Cannostomites

G. †Octomedusa Johnson and Richardson, Jr. 1968

†Octomedusa pieckorum Johnson and Richardson, Jr. 1968

Fm. Periphyllidae Claus 1886

Subfm. Paraphyllininae Maas 1903

G. †Epiphyllina

Fm. †Corumbellidae Hahn et al. 1982

G. †Corumbella

†Corumbella werneri Hahn et al. 1982

Subcl. †Discomedusae Haeckel 1880

Or. †Rhizostomeae

Fm. Leptobrachiidae Claus 1883

G. †Leptobrachites

G. †Marejellia Žalohar et al. 2025

†Marejellia nereis Žalohar et al. 2025

G. †Prothysanostoma Ossian 1973

†Prothysanostoma eleanorae Ossian 1973

G. †Simplicibrachia

Or. Semaeostomeae Agassiz 1862

Fm. Cyaneidae Agassiz 1862

G. Cyanea Peron and Lesueur 1809

Fm. Eulithotidae Kieslinger 1939

G. †Eulithota

Fm. Pelagiidae Gegenbaur 1856

G. Pelagia Peron and Lesueur 1809

Fm. †Semaeostomitidae Harrington and Moore 1956

G. †Semaeostomites

Fm. Ulmariidae Haeckel 1880

G. Discomedusa Clauss 1877

G. †Garraconularia

G. †Hexangulaconularia

†Hexangulaconularia irregularis He 1984

G. †Holoconularia

Or. †Lithorhizotomatida von Ammon 1886

Fm. †Rhizostomitidae Harrington et al. 1956

G. †Rhizostomites Haeckel 1866

Fm. †Olivooidae Steiner 2014

G. †Qinscyphus Liu et al. 2017

†Qinscyphus necopinus Liu et al. 2017

Unr. †Paleoscyphozoa Van Iten et al. 2014

Unr. †Conulatae Moore and Harrington 1956

Or. †Conulariida Miller and Gurley 1896

Subor. †Conchopeltina Moore and Harrington 1956

Fm. †Conchopeltidae Moore and Harrington 1956

Subor. †Conulariina Miller and Gurley 1896

Fm. †Conulariellidae Kiderlen 1937

Fm. †Conulariidae Walcott 1886

Fm. †Conulariopsidae Sugiyama 1942

G. †Ilankirus Sarsembaev and Marusin 2022

†Ilankirus kessyusensis Sarsembaev and Marusin 2022

G. †Vendoconularia Ivantsov and Fedonkin 2002

†Vendoconularia triradiata Ivantsov and Fedonkin 2002

Or. †Conularina Miller and Gurley 1896

Fm. †Carinachitidae He 1987

G. †Carinachites Qian 1977

Invalid names: Mabianoconullus [synonym]

G. †Emeiconularia Qian et al. 1997

Fm. †Hexangulaconulariidae He 1987

G. †Arthrochites Chen 1982

Invalid names: Primaconulariella [synonym], Sacciella He 1984 [synonym]

G. †Hexaconularia He and Yang 1986

G. †Teresconularia Leme et al. 2003

†Teresconularia argentinensis Leme et al. 2003

Or. Coronatae Dong et al. 2013

G. †Olivooides Qian 1977

†Olivooides multisulcatus Qian 1977

G. †Reticulaconularia

G. †Tasmanoconularia

G. †Tirasiana

†Tirasiana cocarda Becker 1985

Invalid names: Lithorhizostomatida [empty], Scyphomedusae Lankester 1881 [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Vision:

blindc

Created:

2010-02-15 05:15:41

Modified:

2010-02-15 07:15:41

Source: c = class

Reference: Aberhan et al. 2004

Age range

Maximum range based only on fossils: base of the Wonokan to the top of the Sarmatian or 635.00000 to 11.63000 Ma
Minimum age of oldest fossil (stem group age): 538.8 Ma

Collections (733 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Ediacaran	635.0 - 538.8	China (Shaanxi)	Anabarites sp. (200480)
Ediacaran	635.0 - 538.8	Russian Federation	Bjarmia cycloplerusa (242127) Vendoconularia triradiata (10759)
Ediacaran	635.0 - 538.8	Mexico (Sonora)	Cambrotubulus decurvatus (10580 10581)
Ediacaran	635.0 - 538.8	Brazil	Corumbella werneri (51753 237528 237535) Paraconularia sp., Corumbella werneri (237533)
Ediacaran	635.0 - 538.8	Norway	Beltanella sp. (51870)
Ediacaran	635.0 - 538.8	USA (Nevada)	Corumbella sp. (7180 231412)
Ediacaran	635.0 - 538.8	Namibia	Corumbella werneri (236924)
Ediacaran	635.0 - 538.8	USA (California)	Beltanella sp. (231437)
Ediacaran	635.0 - 538.8	Russian Federation (Sakha)	Cambrotubulus sp. (99227)
Ediacaran	635.0 - 538.8	Norway (Finnmark Fylke)	Beltanella sp. (10248)
Ediacaran	635.0 - 538.8	Iran	Corumbella sp. (195101 195106 195107)
Ediacaran	635.0 - 538.8	India	Tirasiana sp. (209000) Tirasiana sp., Beltanella sp. (99446 99448)
Ediacaran	635.0 - 538.8	Brazil (Mato Grosso do Sul)	Corumbella werneri (237540 237543)
Ediacaran	635.0 - 538.8	Paraguay	Corumbella werneri (131398 237510)
Vendian	635.0 - 538.8	Brazil	Corumbella werneri (97562)
Wonokan - Poundian	635.0 - 538.8	Canada (Yukon)	Tirasiana sp. (10356 10358)
Jinnigian	538.8 - 530.7	China (Yunnan)	Anabarites trisulcatus (201536 201540) Anabarites trisulcatus, Cambrotubulus decurvatus (201525 201526 201535) Anabarites trisulcatus, Cambrotubulus decurvatus, Arthrochites emeishanensis, Olivooides multisulcatus (201524) Anabarites trisulcatus, Cambrotubulus decurvatus, Hexaconularia sichuanensis, Olivooides multisulcatus (201533 201537 201538 201539) Anabarites trisulcatus, Cambrotubulus decurvatus, Olivooides multisulcatus (201490) Anabarites trisulcatus, Olivooides multisulcatus (201491 201510) Cambrotubulus decurvatus, Olivooides multisulcatus (201532 201534)
Manykaian	538.8 - 529.0	Norway	Tirasiana sp. (51872)
Manykaian	538.8 - 529.0	China (Sichuan)	Carinachites tetrasulcatus, Carinachites curvatornatus, Arthrochites emeishanensis (7818)
Manykaian	538.8 - 529.0	China (Shaanxi)	Hexaconularia sichuanensis (7819)