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Testudo lohanica
Discussion
The species name lohanica refers to the Lohan River, which flows near the fossil site from which the new species comes.
Taxonomy
Testudo lohanica was named by Perez-Garcia et al. (2022). Its type specimen is VPMNS C5659, a partial skeleton (a partial skeleton including the shell (Figs 2F, 3F, 4U–X, 5), appendicular remains (Fig. 5), a partial skull (Figs 5–8), the right lower jaw (Figs 6, 9A–F) and), and it is a 3D body fossil. Its type locality is Crețești-1, which is in a Tortonian floodplain mudstone/sandstone in Romania.
Sister species lacking formal opinion data
Synonymy list
| Year | Name and author |
|---|---|
| 2022 | Testudo lohanica Perez-Garcia et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Testudo lohanica Perez-Garcia et al. 2022
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Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| A. Perez-Garcia et al. 2022 | Member of the genus Testudo characterized by the following autapomorphies: a very strongly curved inward posterior carapace margin, affecting the medial half of the last pair of peripherals and, especially, the pygal plate; diagnostic morphology of the posterior plastral lobe, with the lateral margins of its longer-than-wide xiphiplastra being sub-straight and almost perpendicular to each other from the inguinal notches to near the contact with the femoral-anal sulci; and anterior limbs that are covered distally by many osteoderms that form a continuous protective shield of polygonal elements. It displays the following unique character combination within this genus: shell size >20 cm; maximum carapace width reached approximately in the middle region of its length, and reaching about 75–80% of its maximum length; absence of carapace humps; absence of nuchal notch; slightly wider than long nuchal; usually eight neurals; a generally hexagonal sixth neural; absence of medial contact of the costal series; two suprapygals, with a trapezoidally shaped first suprapygal embracing a lenticular-shaped second suprapygal; lateral margins of the pygal that converge posteriorly; very narrow cervical, lacking a protruding anterior margin; wider than long vertebrals; maximum width of the first vertebral located in its anterior half; posterior sulcus of the fourth vertebral generally on the eighth neural; absence of contact between the fifth vertebral and the tenth pair of marginals; absence or short overlap of the first pair of pleurals on the nuchal; complete overlap of the pleural-marginal sulci on the costal-peripheral sutures; supracaudal scute present; anal notch three times as wide as long; acute angle between both gular-humeral sulci, approximately 75–85°; gulars as long as the humerals in the medial plane or even longer; medial region of the humeral-pectoral sulci overlapping the posterior margin of the entoplastron or very close to it; medial length of the pectorals equivalent to half of that of the hyoplastra or greater; anals medially longer than the femorals; anteromedially directed femoral-anal sulci, defining a posterior angle <110°; inferior processes of the parietals contacting the anterior portion of the quadrates; presence of a medial maxillary ridge; tubercles on the labial ridge of the maxillae; vomer contacting neither the maxillae nor the basisphenoid, its anterior region being fan-shaped; small foramina palatinum posterius; canalis pro ramo nervi vidiani not contacting the canalis caroticus internus; presence of a branch of the Vidian nerve posterior to the canalis pro ramo nervi vidiani. |
Measurements
No measurements are available
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| Source: f = family, c = class, subp = subphylum | |||||
| References: Hendy et al. 2009, Ernst and Barbour 1989, Carroll 1988 | |||||